Contents 1 Regulation 2 See also 3 References 4 External links


Regulation[edit] Plants regulate the rate of transpiration by controlling the size of the stomatal apertures. The rate of transpiration is also influenced by the evaporative demand of the atmosphere surrounding the leaf such as boundary layer conductance, humidity, temperature, wind and incident sunlight. Soil water supply and soil temperature can influence stomatal opening, and thus transpiration rate. The amount of water lost by a plant also depends on its size and the amount of water absorbed at the roots. Transpiration accounts for most of the water loss by a plant by the leaves and young stems. Transpiration serves to evaporatively cool plants, as the evaporating water carries away heat energy due to its large latent heat of vaporization of 2260 kJ per litre. Feature Effect on transpiration Number of leaves More leaves (or spines, or other photosynthesizing organs) means a bigger surface area and more stomata for gaseous exchange. This will result in greater water loss. Number of stomata More stomata will provide more pores for transpiration. Size of the leaf A leaf with a bigger surface area will transpire faster than a leaf with a smaller surface area. Presence of plant cuticle A waxy cuticle is relatively impermeable to water and water vapour and reduces evaporation from the plant surface except via the stomata. A reflective cuticle will reduce solar heating and temperature rise of the leaf,[citation needed] helping to reduce the rate of evaporation. Tiny hair-like structures called trichomes on the surface of leaves also can inhibit water loss by creating a high humidity environment at the surface of leaves.[citation needed] These are some examples of the adaptations of plants for conservation of water that may be found on many xerophytes. Light supply The rate of transpiration is controlled by stomatal aperture, and these small pores open especially for photosynthesis. While there are exceptions to this (such as night or "CAM photosynthesis"), in general a light supply will encourage open stomata. Temperature Temperature affects the rate in two ways: 1) An increased rate of evaporation due to a temperature rise will hasten the loss of water. 2) Decreased relative humidity outside the leaf will increase the water potential gradient. Relative humidity Drier surroundings gives a steeper water potential gradient, and so increases the rate of transpiration. Wind In still air, water lost due to transpiration can accumulate in the form of vapor close to the leaf surface. This will reduce the rate of water loss, as the water potential gradient from inside to outside of the leaf is then slightly less. Wind blows away much of this water vapor near the leaf surface, making the potential gradient steeper and speeding up the diffusion of water molecules into the surrounding air. Even in wind, though, there may be some accumulation of water vapor in a thin boundary layer of slower moving air next to the leaf surface. The stronger the wind, the thinner this layer will tend to be, and the steeper the water potential gradient. Water supply Water stress caused by restricted water supply from the soil may result in stomatal closure and reduce the rates of transpiration. The effect of temperature on the transpiration rate of plants. The effect of wind velocity on the transpiration rate of plants. The effect of humidity on the transpiration rate of plants. Some xerophytes will reduce the surface of their leaves during water deficiencies (left). If temperatures are cool enough and water levels are adequate the leaves expand again (right). During a growing season, a leaf will transpire many times more water than its own weight. An acre of corn gives off about 3,000–4,000 gallons (11,400–15,100 liters) of water each day, and a large oak tree can transpire 40,000 gallons (151,000 liters) per year. The transpiration ratio is the ratio of the mass of water transpired to the mass of dry matter produced; the transpiration ratio of crops tends to fall between 200 and 1000 (i.e., crop plants transpire 200 to 1000 kg of water for every kg of dry matter produced).[6] Transpiration rates of plants can be measured by a number of techniques, including potometers, lysimeters, porometers, photosynthesis systems and thermometric sap flow sensors. Isotope measurements indicate transpiration is the larger component of evapotranspiration.[7] Recent evidence from a global study[8] of water stable isotopes shows that transpired water is isotopically different from groundwater and streams. This suggests that soil water is not as well mixed as widely assumed.[9] Desert plants have specially adapted structures, such as thick cuticles, reduced leaf areas, sunken stomata and hairs to reduce transpiration and conserve water. Many cacti conduct photosynthesis in succulent stems, rather than leaves, so the surface area of the shoot is very low. Many desert plants have a special type of photosynthesis, termed crassulacean acid metabolism or CAM photosynthesis, in which the stomata are closed during the day and open at night when transpiration will be lower.


See also[edit] Antitranspirant – a substance to prevent transpiration Eddy covariance flux (aka eddy correlation, eddy flux) Hydrology (agriculture) Latent heat flux Perspiration Soil plant atmosphere continuum Water Evaluation And Planning system (WEAP) Transpiration stream Wikisource has the text of the 1905 New International Encyclopedia article Transpiration.


References[edit] ^ Sinha, Rajiv Kumar (2004-01-01). Modern Plant Physiology. CRC Press. ISBN 978-0-8493-1714-9.  ^ Benjamin Cummins (2007), Biological Science (3 ed.), Freeman, Scott, p. 215  ^ Taiz, Lincoln (2015). Plant Physiology and Development. Sunderland, MA: Sinauer Associates, Inc. p. 101. ISBN 978-1-60535-255-8.  ^ Freeman, Scott (2014). Biological Sciences. United States of America: Pearson. pp. 765–766. ISBN 978-0-321-74367-1.  ^ Graham, Linda E. (2006). Plant Biology. Upper Saddle River, NJ 07458: Pearson Education, Inc. pp. 200–202. ISBN 0-13-146906-1.  ^ Martin, J.; Leonard, W.; Stamp, D. (1976), Principles of Field Crop Production (3rd ed.), New York: Macmillan Publishing Co., ISBN 0-02-376720-0  ^ Jasechko, Scott; Sharp, Zachary D.; Gibson, John J.; Birks, S. Jean; Yi, Yi; Fawcett, Peter J. (3 April 2013). "Terrestrial water fluxes dominated by transpiration". Nature. 496 (7445): 347–50. doi:10.1038/nature11983. PMID 23552893. Retrieved 4 April 2013.  ^ Evaristo, Jaivime; Jasechko, Scott; McDonnell, Jeffrey J. (2015-09-03). "Global separation of plant transpiration from groundwater and streamflow". Nature. NPG. 525 (7567): 91–94. doi:10.1038/nature14983. ISSN 0028-0836. PMID 26333467.  ^ Bowen, Gabriel (2015-09-03). "Hydrology: The diversified economics of soil water". Nature. 525 (7567): 43–44. doi:10.1038/525043a. ISSN 0028-0836. PMID 26333464. 


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