Contents 1 Distribution and habitat 2 Morphology 3 Taxonomy 4 Behavior 4.1 Vision 4.2 Mating 4.3 Sexual selection 4.3.1 Sexual selection experiments 4.4 Handicap selection 5 References 6 External links

Distribution and habitat[edit] More than 100 species in the Diopsidae are known, with the greatest diversity found in the Old World tropics.[2] They are distributed throughout the region, with the best-known species being from Southeast Asia and Southern Africa. Also, two species in North America have been described and a European species has recently been found in Hungary.[3] Adult diopsids are typically found on low-lying vegetation in humid areas, often near streams and rivers, where they feed on fungi and bacteria, which they scavenge from decaying vegetation. The larvae are saprophagic or phytophagous, eating decaying and fresh plant matter. Diopsis macrophthalma Dalman, 1817, is a pest of rice and sorghum in tropical Africa. The peculiar morphology of stalk-eyed flies makes it easy to identify their fossils (e.g. in amber); one prehistoric genus that has been discovered in that form is Prosphyracephala.[4]

Morphology[edit] The Diopsidae are small to medium-sized flies, ranging from about 4.0 to about 12.0 mm in length. Their heads are subtriangular, with transverse eye stalks in all genera except the African genus Centrioncus. The head is usually sparsely haired, with vibrissae (whiskers) absent.[5] The posterior portion of the fly's metathorax, or scutellum, has a pair of stout processes, and often the laterotergite (one of a number of lateral flanges) of the postnotum (a small dorsal sclerite on the insect thorax posterior to the notum) has a dome-like swelling or spine-like process. The anterior femora of the legs are stout, with ventral spines. Adult males have lost tergites seven and eight, and the seventh sternite forms a complete ventral band.[5] Stalk-eyed flies, as the name implies, typically possess eyestalks. Their eyes are mounted on projections from the sides of the head, and the antennae are located on the eyestalks, unlike stalk-eyed flies from other families. A rather remarkable feature of stalk-eyed flies is their ability, shortly after they emerge from their pupae, to ingest air through their oral cavity and pump it through ducts in the head to the tips of the eye stalks, thereby elongating them while they are still soft and transparent.[6]

Taxonomy[edit] Main article: List of stalk-eyed flies True stalk-eyed flies are members of the family Diopsidae, first described by Fothergill[7] and named by Carl Linnaeus in 1775.[8] The family Diopsidae is contained within the order Diptera and suborder Cyclorrapha, and features some 150 described species.[9] The African genus Centrioncus (once placed in Sepsidae, but then moved to Diopsidae) was once recommended to be treated as a separate family, Centrioncidae, a sister group of the diopsids,[10] but since then this lineage has usually been treated as a subfamily.[11]

Behavior[edit] Vision[edit] Despite the unusual morphology of the eye, each compound eye sees a region of space extending over more than a hemisphere in all directions. Thus, extensive binocular overlap occurs, with about 70% of the ommatidia of each eye having a binocular partner ommatidium in the opposite eye which views in the same direction.[12][13] The binocular field is most extensive in the frontoventral quadrant, where it reaches over 135°, and is smallest in the dorsal region. The behavior of stalk-eyed flies is very much determined by vision. During the day, temporary territories may be defended by threatening behavior. At dusk, the animals gather in small groups on selected thread-like structures, returning to the same site each day. When males of about equal size encounter one another within such a group, they may engage in ritualized fights (or occasionally contact fights). Competitors are driven away by the dominant male. Conspecifics are most likely to elicit a threat or flight reaction when they are at a distance of about 50 mm, and reactions to model flies and reflections in a mirror also occur at about this distance.[12] Mating[edit] Male Teleopsis dalmanni Stalk-eyed flies roost at night on root hairs hanging by streams. Mating usually takes place in the early morning in the vicinity of their roosts. Females show a strong preference for roosting and mating with males with longer eyestalks, and males compete with each other to control lekking aggregations through ritualized contest. This contest involves males facing one another and comparing their relative eye spans, often with the front legs spread apart, possibly to emphasize their eye-stalk lengths .[14] Male stalk-eyed flies with long eyestalks gain mating advantages both because of female choice and because they are better able to compete with rival males.[2][15] Sexual selection[edit] Though the evolution of exaggerated male traits as a result of female mate choice was at one point controversial, the Diopsidae are now regarded as a classic example of animals that exhibit sexually selected traits.[16][17][18] One view maintains that male ornaments co-evolve with female preferences. The selection of an ornamented mate causes genes that influence expression of the selected male trait and genes coding for female preference for this trait to be passed on to offspring[19][20] This process creates linkage disequilibrium between selected alleles, with the magnitude of resulting genetic correlations influencing evolutionary outcomes. If the genetic correlation is high relative to the heritability of the male ornament, then a runaway process can occur leading to extreme sexually selected traits, such as the incredible eye spans observed in male stalk-eyed flies. Otherwise, the trait and preference for the trait increase until viability natural selection against further trait elaboration balances sexual selection.[19] Close-up of a male Teleopsis dalmanni The extreme morphology exhibited by stalk-eyed flies (especially males) has been studied in an effort to support the hypothesis that exaggerated male traits could evolve through female mate choice and that the selection on male ornaments should cause a correlated response in female preferences. Researchers noted that the flies roosted along stream banks in peninsular Malaysia and that the males with the largest eye spans were accompanied by more females than males with shorter eye spans. From January to October, the researchers counted males and females on 40 root hairs along a single 200-m stretch of stream bank to confirm this observation.[21] Sexual selection experiments[edit] Researchers collected stalk-eyed flies and observed their behavior under laboratory conditions. In the lab, each individual was scored for eye spans, body length, age, and fecundity.[22] Pairs of males differing in eye span but matched in body length were conducted to quantify mate choice in the presence and absence of male interactions. Test males with the longest or shortest eye-span to body-length ratios were mated with 25 randomly chosen females. Wilkinson and Reillo then tested female choice in the presence and absence of male competition and in the presence of males with abnormally long and abnormally short eye spans.[21] Males dispersed themselves, while females clustered in certain areas of the cage. As observed prior to the study, researchers found that the average number of females per male increased with male eye span in field collected aggregations of stalk-eyed flies. Under laboratory conditions, researchers found that female preferences for male characteristics changed as the males sexual characteristics changed. After 13 generations of artificial selection, they found that long eye-span male line females (i.e. females whose fathers had long eye spans) preferred long eye spans in both the selected males and in males that were not bred through artificial selection, while short eye-span male line females (i.e. females whose fathers had short eye spans) found short eye spans to be the most attractive, even over males with long eye spans. Because researchers kept the females separate from males prior to mate selection, the finding supported the hypothesis that the change in female mate choice was genetically based and not learned. Thus, stalk-eyed flies have been able to evolve sexual trait in males that corresponds directly to traits that affect mating choices made by females.[21] Stalk-eyed fly (Diasemopsis) Handicap selection[edit] However, the evolution of extreme morphology in male flies and the corresponding evolution of female preference for these characteristics as an effect of sexual selection is only half the picture.[23] Handicap models of sexual selection predict that male sexual ornaments have strong condition-dependent expression, and this allows females to evaluate male genetic quality.[24][25][26][27][28] Genetic variation underlies the response to environmental stress, such as variable food quality, of male sexual ornaments, such as the increased eye span, in the stalk-eyed fly.[23] Some male genotypes develop large eye spans under all conditions, whereas other genotypes progressively reduce eye spans as environmental conditions deteriorate. Several nonsexual traits, including female eye span and male and female wing length, also show condition-dependent expression, but their genetic response is entirely explained by scaling with body size. Unlike these characteristics, male eye span still reveals genetic variation in response to environmental stress after accounting for differences in body size. Thus, it could be inferred that these results strongly support the conclusion that female mate choice yields genetic benefits for offspring as eye span acts as a truthful indicator of male fitness. Eye span is, therefore, selected not only on the basis of attractiveness, but also because it demonstrates good genes in mates.[23] Furthermore, some populations of stalk-eyed fly females carry a meiotic drive gene on their X chromosomes that causes female-biased sex ratios.[29] In these populations, males which carry a gene to suppresses X chromosome meiotic drive have longer eyestalks. Thus, females that mate with these males gain a direct genetic benefit by producing male offspring in a female-biased population. In other words, the gene for long eye-stalks is linked to a gene that makes males sire more male offspring.[30] Alternatively, long stalks may signal fertility, perhaps by encouraging females to use the sperm of a long-stalked male so as to produce more fertile sons.[30]

References[edit] ^ Feijen, Hans R. (1983). "Systematics and phylogeny of Centrioncidae, a new afromontane family of Diptera (Schizophora)". Zoologische Verhandelingen. Leiden. 202: 1–137. Retrieved 30 October 2016.  ^ a b G. S. Wilkinson & G. N. Dodson (1997). "Function and evolution of antlers and eye stalks in flies". In J. Choe & B. Crespi. The Evolution of Mating Systems in Insects and Arachnids. Cambridge: Cambridge University Press. pp. 310–327. ISBN 0-521-58976-2.  ^ L. Papp, M. Földvári & P. Paulovics (1997). "Sphyracephala europaea sp. n. (Diptera: Diopsidae) from Hungary represents a family new to Europe". Folia Entomologica Hungarica. 58: 137–146.  ^ Schumann, H (1994). "Diopsidae from Saxon Amber (Diptera, Diopsidae)". Deutsche Entomologische Zeitschrift. 41 (1): 141–145. doi:10.1002/mmnd.19940410111.  ^ a b Peterson, B. V. (1987). "Diopsidae." In McAlpine, J. F., B. V. Peterson, G. E. Sherwell, H. J. Tekey, J. R. Vockerorth, and D. M. Wood, (cords.). Manual of Nearctic Diptera. Vol. 2: 785–789. ^ Buschbeck, E. K., Roosevelt, J. L. and Hoy, R. R. (2001). Eye stalks or no eye stalks: A structural comparison of pupal development in the stalk-eyed fly Cyrtodiopsis and in Drosophila. J. Comp. Neurol., 433: 486–498. ^ Shillito, J. (1976). Fothergill and Linnaeus: The background of De Bigis Insectorum, 1775. Biol. J. Linn. Soc., 8; 75-86. ^ Shillito, J. (1976). Bibliography of the Diopsidae-II. Journal of the Society for the Bibliography of Natural History. Volume 8, Page 65-73. ^ Allaby, M. (1999). "Diopsidae." A Dictionary of Zoology. (December 15, 2012). ^ Feijen, H.R. (1983). "Systematics and phylogeny of Centrioncidae, a new afromontane family of Diptera (Schizophora)". 202 (1): 1–137.  ^ Meier, R. & Hilger, S. (2000). "On the egg morphology and phylogenetic relationships of Diopsidae (Diptera: Schizophora)". Journal of Zoological Systematics and Evolutionary Research. 38 (1): 1–36. doi:10.1046/j.1439-0469.2000.381128.x.  ^ a b D. Burkhardt & I. de la Motte (1983). "How stalk-eyed flies eye stalk-eyed flies: Observations and measurements of the eyes of Cyrtodiopsis whitei (Diopsidae, Diptera)". Journal of Comparative Physiology A: Neuroethology, Sensory, Neural, and Behavioral Physiology. 151 (4): 407–421. doi:10.1007/BF00605457.  ^ I. de la Motte & D. Burkhardt (1983). "Portrait of an Asian stalk-eyed fly". Naturwissenschaften. 70 (9): 451–461. doi:10.1007/BF01079611.  ^ Wilkinson, G. (2001). Model Syestems in Behavioral Ecology: Integrating Conceptual, Theoretical, and Empirical Approaches. Ed. Lee Alan Dugatkin. Princeton University Press; Princeton: pp. 84-91. ^ T. Chapman, A. Pomiankowski & K. Fowler (2005). "Quick guide: stalk-eyed flies". Current Biology. 15 (14): R533–R535. doi:10.1016/j.cub.2005.07.015. PMID 16051154.  ^ Davies N, Krebs J, and West S. (2012). An Introduction to Behavioral Ecology, 4th Ed. Wiley-Blackwell; Oxford: pp. 196-198. ^ Kirkpatrick, M and M.J. Ryan. (1991). “The evolution of mating preferences and the paradox of the lek.” Nature, Lond. 350: 33-38. ^ Maynard Smith, J. (1991). “Theories of sexual selection.” Trends Ecol. Evol. 6: 146-151. ^ a b Lande, R. (1981). “Models of speciation by sexual selection on polygenic traits.” Proc. Natn. Acad. Sci. U.S.A. 78: 3721-3725. ^ Kirkpatrick, M. (1982). “Sexual selection and the evolution of female choice.” Evolution 36: 1-12. ^ a b c Wilkinson, G. and P. Reillo. (1994). “Female choice response to artificial selection on an exaggerated male trait in a stalk-eyed fly.” Proc. R. Soc. Lond. B. 255: 1-6. ^ Wilkinson, G. (1993). “Artificial sexual selection alters allometry in the stalk-eyed fly Cyrtodiopsis dalmanni.” Genet. Res. 62: 212-222. ^ a b c P. David, T. Bjorksten, K. Fowler & A. Pomiankowski (2000). "Condition-dependent signalling of genetic variation in stalk-eyed flies". Nature. 406 (6792): 186–188. doi:10.1038/35018079. PMID 10910358. CS1 maint: Multiple names: authors list (link) ^ Andersson, M. (1986). “Evolution of condition-dependent sex ornaments and mating preferences: sexual selection based on viability differences.” Evolution 40: 804–816. ^ Pomiankowski, A. (1987). “Sexual selection: the handicap principle does work—sometimes.” Proc. R. Soc. Lond. B 231: 123–145. ^ Grafen, A. (1990). “Biological signals as handicaps.” J. Theor. Biol. 144: 517–546. ^ Iwasa, Y. and A. Pomiankowski. (1994). “The evolution of mate preferences for multiple handicaps.” Evolution 48: 853 –867. ^ Rowe, L. and D. Houle. (1996). “The lek paradox and the capture of genetic variance by condition dependent traits.” Proc. R. Soc. Lond. B 263: 1415–1421. ^ G. S. Wilkinson, D. C. Presgraves &. L. Crymes (1998). "Male eye span in stalk-eyed flies indicates genetic quality by meiotic drive suppression". Nature. 391 (6664): 276–279. doi:10.1038/34640.  ^ a b Zimmer, Carl (2008). "The Evolution of Extraordinary Eyes: The Cases of Flatfishes and Stalk-eyed Flies". Evolution Education and Outreach. 1 (4): 487. doi:10.1007/s12052-008-0089-9. 

External links[edit] Wikispecies has information related to Diopsidae Wikimedia Commons has media related to Diopsidae. Family Diopsidae at EOL Stalk-eyed Fly Research Group Gerald Wilkinson's lab Pest Information Wiki v t e Extant Diptera families Kingdom: Animalia Phylum: Arthropoda Class: Insecta Subclass: Pterygota Infraclass: Neoptera Superorder: Endopterygota Suborder Nematocera Axymyiomorpha Axymyiidae Culicomorpha Culicoidea Dixidae (meniscus midges) Corethrellidae (frog-biting midges) Chaoboridae (phantom midges) Culicidae (mosquitoes) Chironomoidea Thaumaleidae (solitary midges) Simuliidae (black flies) Ceratopogonidae (biting midges) Chironomidae (non-biting midges) Blephariceromorpha Blephariceridae (net-winged midges) Deuterophlebiidae (mountain midges) Nymphomyiidae Bibionomorpha Bibionoidea Bibionidae (march flies, lovebugs) Anisopodoidea Anisopodidae (wood gnats) Sciaroidea (fungus gnats) Bolitophilidae Diadocidiidae Ditomyiidae Keroplatidae Mycetophilidae Sciaridae (dark-winged fungus gnats) Cecidomyiidae (gall midges) Psychodomorpha Scatopsoidea Canthyloscelidae Perissommatidae Scatopsidae (minute black scavenger flies, or dung midges) Psychodoidea Psychodidae (moth flies) Ptychopteromorpha Ptychopteridae (phantom crane flies) Tanyderidae (primitive crane flies) Tipulomorpha Trichoceroidea Trichoceridae (winter crane flies) Tipuloidea Pediciidae (hairy-eyed craneflies) Tipulidae (crane flies) Suborder Brachycera Asilomorpha Asiloidea Apioceridae (flower-loving flies) Apsilocephalidae Apystomyiidae Asilidae (robber flies) Bombyliidae (bee flies) Evocoidae Hilarimorphidae (hilarimorphid flies) Mydidae (mydas flies) Mythicomyiidae Scenopinidae (window flies) Therevidae (stiletto flies) Empidoidea Atelestidae Hybotidae (dance flies) Dolichopodidae (long-legged flies) Empididae (dagger flies, balloon flies) Nemestrinoidea Acroceridae (small-headed flies) Nemestrinidae (tangle-veined flies) Muscomorpha Aschiza Platypezoidea Phoridae (scuttle flies, coffin flies, humpbacked flies) Opetiidae (flat-footed flies) Ironomyiidae (ironic flies) Lonchopteridae (spear-winged flies) Platypezidae (flat-footed flies) Syrphoidea Syrphidae (hoverflies) Pipunculidae (big-headed flies) Schizophora Acalyptratae Conopoidea Conopidae (thick-headed flies) Tephritoidea Pallopteridae (flutter flies) Piophilidae (cheese flies) Platystomatidae (signal flies) Pyrgotidae Richardiidae Tephritidae (peacock flies) Ulidiidae (picture-winged flies) Nerioidea Cypselosomatidae Micropezidae (stilt-legged flies) Neriidae (cactus flies, banana stalk flies) Pseudopomyzidae (banana flies) Diopsoidea Diopsidae (stalk-eyed flies) Gobryidae Megamerinidae Nothybidae Psilidae (rust flies) Somatiidae Strongylophthalmyiidae Syringogastridae Tanypezidae Sciomyzoidea Coelopidae (kelp flies) Dryomyzidae Helosciomyzidae Ropalomeridae Huttoninidae Heterocheilidae Phaeomyiidae Sepsidae (black scavenger flies) Sciomyzidae (marsh flies) Sphaeroceroidea Chyromyidae Heleomyzidae Sphaeroceridae (small dung flies) Nannodastiidae Lauxanioidea Celyphidae (beetle-backed flies) Chamaemyiidae (aphid flies) Lauxaniidae Opomyzoidea Agromyzidae (leaf miner flies) Anthomyzidae Asteiidae Aulacigastridae (sap flies) Clusiidae (lekking, or druid flies) Fergusoninidae Marginidae Neminidae Neurochaetidae (upside-down flies) Odiniidae Opomyzidae Periscelididae Teratomyzidae Xenasteiidae Ephydroidea Camillidae Curtonotidae (quasimodo flies) Diastatidae (bog flies) Ephydridae (shore flies) Drosophilidae (vinegar and fruit flies) Carnoidea Acartophthalmidae Australimyzidae Braulidae (bee lice) Canacidae (beach flies) Carnidae Chloropidae (frit flies) Cryptochaetidae Inbiomyiidae Milichiidae (freeloader flies) Lonchaeoidea Cryptochetidae Lonchaeidae (lance flies) Calyptratae Muscoidea Anthomyiidae (cabbage flies) Fanniidae (little house flies) Muscidae (house flies, stable flies) Scathophagidae (dung flies) Oestroidea Calliphoridae (blow-flies: bluebottles, greenbottles) Mystacinobiidae (New Zealand bat fly) Oestridae (botflies) Rhinophoridae Sarcophagidae (flesh flies) Tachinidae (tachina flies) Hippoboscoidea Glossinidae (tsetse flies) Hippoboscidae (louse flies) Mormotomyiidae (frightful hairy fly) Nycteribiidae (bat flies) Streblidae (bat flies) Stratiomyomorpha Stratiomyoidea Pantophthalmidae (timber flies) Stratiomyidae (soldier flies) Xylomyidae (wood soldier flies) Tabanomorpha Rhagionoidea Austroleptidae Bolbomyiidae Rhagionidae (snipe flies) Tabanoidea Athericidae (water snipe flies) Oreoleptidae Pelecorhynchidae Tabanidae (horse and deer flies) Vermileonomorpha Vermileonoidea Vermileonidae Xylophagomorpha Xylophagoidea Xylophagidae (awl flies) List of families of Diptera Retrieved from "" Categories: DiopsidaeHidden categories: CS1 maint: Multiple names: authors listArticles with 'species' microformats

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Stalk-eyed_fly - Photos and All Basic Informations

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CameroonTaxonomy (biology)AnimalArthropodInsectFlyCyclorrhaphaMuscomorphaSchizophoraAcalyptrataeDiopsoideaGenusCentrioncusCladodiopsisDiasemopsisDiopsinaEosiopsisEurydiopsisPseudodiopsisSphyracephalaTeleopsisTeloglabrusSynonym (taxonomy)FlyDrosophilidaePlatystomatidaeRichardiidaeTephritidaeAntenna (biology)Morphology (biology)Sexual SelectionNatural SelectionSexual SelectionHandicap PrincipleOld WorldSoutheast AsiaSouthern AfricaNorth AmericaSaprophagousPhytophagousRiceSorghumFossilAmberProsphyracephalaCentrioncusVibrissaeMetathoraxScutellum (insect)ScleriteNotumTergitesSterniteVentralList Of Stalk-eyed FliesJohn Fothergill (physician)Carl LinnaeusDipteraCentrioncusSepsidaeCompound EyeOmmatidiaConspecificsEnlargeLek (mating Arena)Linkage DisequilibriumAllelesHeritabilityNatural SelectionSexual SelectionEnlargeMalaysiaFecundityFemale ChoiceEnlargeDiasemopsisHandicap PrincipleFitness (biology)Meiotic DriveSex RatioCambridge University PressInternational Standard Book NumberSpecial:BookSources/0-521-58976-2Digital Object IdentifierDigital Object IdentifierDigital Object IdentifierNaturwissenschaftenDigital Object IdentifierCurrent BiologyDigital Object IdentifierPubMed IdentifierNature (journal)Digital Object IdentifierPubMed IdentifierCategory:CS1 Maint: Multiple Names: Authors ListNature (journal)Digital Object IdentifierDigital Object IdentifierWikispeciesTemplate:DipteraTemplate Talk:DipteraFlyAnimalArthropodInsectPterygotaNeopteraEndopterygotaNematoceraAxymyiidaeAxymyiidaeCulicomorphaCulicoideaDixidaeCorethrellidaeChaoboridaeMosquitoChironomoideaThaumaleidaeBlack FlyCeratopogonidaeChironomidaeBlephariceromorphaBlephariceridaeDeuterophlebiaNymphomyiidaeBibionomorphaBibionidaeBibionidaeAnisopodidaeAnisopodidaeSciaroideaBolitophilaDiadocidiidaeDitomyiidaeKeroplatidaeMycetophilidaeSciaridaeCecidomyiidaePsychodomorphaScatopsoideaCanthyloscelidaePerissommatidaeScatopsidaePsychodidaePsychodidaePtychopteromorphaPtychopteridaeTanyderidaeTipulomorphaTrichoceridaeTrichoceridaeTipuloideaPediciidaeCrane FlyBrachyceraAsilomorphaAsiloideaApioceraApsilocephalidaeApystomyiidaeAsilidaeBombyliidaeEvocoaHilarimorphidaeMydas FlyMythicomyiidaeScenopinidaeTherevidaeEmpidoideaAtelestidaeHybotidaeDolichopodidaeEmpididaeNemestrinoideaAcroceridaeNemestrinidaeMuscomorphaAschizaPlatypezoideaPhoridaeOpetiidaeIronomyiidaeLonchopteridaePlatypezidaeSyrphoideaHoverflyPipunculidaeSchizophoraAcalyptrataeConopidaeConopidaeTephritoideaPallopteridaePiophilidaePlatystomatidaePyrgotidaeRichardiidaeTephritidaeUlidiidaeNerioideaCypselosomatidaeMicropezidaeNeriidaePseudopomyzidaeDiopsoideaGobryidaeMegamerinidaeNothybidaePsilidaeStrongylophthalmyiidaeSyringogasterTanypezidaeSciomyzoideaCoelopidaeDryomyzidaeHelosciomyzidaeRopalomeridaeHuttoninidaeHeterocheilidaePhaeomyiidaeSepsidaeSciomyzidaeSphaeroceroideaChyromyidaeHeleomyzidaeSphaeroceridaeLauxanioideaCelyphidaeChamaemyiidaeLauxaniidaeOpomyzoideaAgromyzidaeAnthomyzidaeAsteiidaeAulacigastridaeClusiidaeOdiniidaeOpomyzidaePeriscelididaeEphydroideaCamillidaeCurtonotidaeDiastatidaeEphydridaeDrosophilidaeCarnoideaAcartophthalmusAustralimyzidaeBraulidaeCanacidaeCarnidaeChloropidaeMilichiidaeCryptochetidaeLonchaeidaeCalyptrataeMuscoideaAnthomyiidaeFanniidaeMuscidaeScathophagidaeOestroideaCalliphoridaeNew Zealand Bat FlyBotflyRhinophoridaeFlesh-flyTachinidaeHippoboscoideaTsetse FlyHippoboscidaeMormotomyiidaeNycteribiidaeStreblidaeStratiomyomorphaStratiomyoideaPantophthalmidaeStratiomyidaeXylomyidaeTabanomorphaAustroleptidaeBolbomyiidaeRhagionidaeTabanoideaAthericidaePelecorhynchidaeHorse-flyVermileonidaeVermileonidaeVermileonidaeXylophagidaeXylophagidaeXylophagidaeList Of Families Of DipteraHelp:CategoryCategory:DiopsidaeCategory:CS1 Maint: Multiple Names: Authors ListCategory:Articles With 'species' MicroformatsDiscussion About Edits From This IP Address [n]A List Of Edits Made From This IP Address [y]View The Content Page [c]Discussion About The Content Page [t]Edit This Page [e]Visit The Main Page [z]Guides To Browsing WikipediaFeatured Content – The Best Of WikipediaFind Background Information On Current EventsLoad A Random Article [x]Guidance On How To Use And Edit WikipediaFind Out About WikipediaAbout The Project, What You Can Do, Where To Find ThingsA List Of Recent Changes In The Wiki [r]List Of All English Wikipedia Pages Containing Links To This Page [j]Recent Changes In Pages Linked From This Page [k]Upload Files [u]A List Of All Special Pages [q]Wikipedia:AboutWikipedia:General Disclaimer

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