Contents 1 Definition 2 History 3 Theory 4 Support for premise 5 Reception 6 See also 7 Notes 8 References 8.1 Bibliography 9 External links


Definition[edit] E. O. Wilson defined sociobiology as "the extension of population biology and evolutionary theory to social organization".[1] Sociobiology is based on the premise that some behaviors (social and individual) are at least partly inherited and can be affected by natural selection. It begins with the idea that behaviors have evolved over time, similar to the way that physical traits are thought to have evolved. It predicts that animals will act in ways that have proven to be evolutionarily successful over time. This can, among other things, result in the formation of complex social processes conducive to evolutionary fitness. The discipline seeks to explain behavior as a product of natural selection. Behavior is therefore seen as an effort to preserve one's genes in the population. Inherent in sociobiological reasoning is the idea that certain genes or gene combinations that influence particular behavioral traits can be inherited from generation to generation[2] For example, newly dominant male lions often kill cubs in the pride that they did not sire. This behavior is adaptive because killing the cubs eliminates competition for their own offspring and causes the nursing females to come into heat faster, thus allowing more of his genes to enter into the population. Sociobiologists would view this instinctual cub-killing behavior as being inherited through the genes of successfully reproducing male lions, whereas non-killing behavior may have died out as those lions were less successful in reproducing.[3]


History[edit] E. O. Wilson, a central figure in the history of sociobiology, from the publication in 1975 of his book Sociobiology: The New Synthesis The ethologist John Paul Scott coined the word sociobiology at a 1948 conference on genetics and social behaviour,[4] and it became widely used after it was popularized by Edward O. Wilson in his 1975 book, Sociobiology: The New Synthesis. However, the influence of evolution on behavior has been of interest to biologists and philosophers since soon after the discovery of evolution itself. Peter Kropotkin's Mutual Aid: A Factor of Evolution, written in the early 1890s, is a popular example. Antecedents of modern sociobiological thinking can be traced to the 1960s and the work of such biologists as Richard D. Alexander, Robert Trivers and William D. Hamilton. The idea of the inheritance of behaviour arose from J. B. S. Haldane's idea about how "altruistic behaviour" (see Altruism) could be passed from generation to generation.[5] Wilson's book pioneered and popularized the attempt to explain the evolutionary mechanics behind social behaviors such as altruism, aggression, and nurturance, primarily in ants (Wilson's own research specialty) and other Hymenoptera, but also in other animals.[6] The final chapter of the book is devoted to sociobiological explanations of human behavior, and Wilson later wrote a Pulitzer Prize winning book, On Human Nature, that addressed human behavior specifically.[7] Edward H. Hagen writes in The Handbook of Evolutionary Psychology that sociobiology is, despite the public controversy regarding the applications to humans, "one of the scientific triumphs of the twentieth century." "Sociobiology is now part of the core research and curriculum of virtually all biology departments, and it is a foundation of the work of almost all field biologists" Sociobiological research on nonhuman organisms has increased dramatically and continuously in the world's top scientific journals such as Nature and Science. The more general term behavioral ecology is commonly substituted for the term sociobiology in order to avoid the public controversy.[8]


Theory[edit] Sociobiologists believe that human behavior, as well as nonhuman animal behavior, can be partly explained as the outcome of natural selection. They contend that in order to fully understand behavior, it must be analyzed in terms of evolutionary considerations. Natural selection is fundamental to evolutionary theory. Variants of hereditary traits which increase an organism's ability to survive and reproduce will be more greatly represented in subsequent generations, i.e., they will be "selected for". Thus, inherited behavioral mechanisms that allowed an organism a greater chance of surviving and/or reproducing in the past are more likely to survive in present organisms. That inherited adaptive behaviors are present in nonhuman animal species has been multiply demonstrated by biologists, and it has become a foundation of evolutionary biology. However, there is continued resistance by some researchers over the application of evolutionary models to humans, particularly from within the social sciences, where culture has long been assumed to be the predominant driver of behavior. Nikolaas Tinbergen, whose work influenced sociobiology. Sociobiology is based upon two fundamental premises: Certain behavioral traits are inherited, Inherited behavioral traits have been honed by natural selection. Therefore, these traits were probably "adaptive" in the environment in which the species evolved. Sociobiology uses Nikolaas Tinbergen's four categories of questions and explanations of animal behavior. Two categories are at the species level; two, at the individual level. The species-level categories (often called "ultimate explanations") are the function (i.e., adaptation) that a behavior serves and the evolutionary process (i.e., phylogeny) that resulted in this functionality. The individual-level categories (often called "proximate explanations") are the development of the individual (i.e., ontogeny) and the proximate mechanism (e.g., brain anatomy and hormones). Sociobiologists are interested in how behavior can be explained logically as a result of selective pressures in the history of a species. Thus, they are often interested in instinctive, or intuitive behavior, and in explaining the similarities, rather than the differences, between cultures. For example, mothers within many species of mammals – including humans – are very protective of their offspring. Sociobiologists reason that this protective behavior likely evolved over time because it helped the offspring of the individuals which had the characteristic to survive. This parental protection would increase in frequency in the population. The social behavior is believed to have evolved in a fashion similar to other types of nonbehavioral adaptations, such as a coat of fur, or the sense of smell. Individual genetic advantage fails to explain certain social behaviors as a result of gene-centred selection. E.O. Wilson argued that evolution may also act upon groups.[9] The mechanisms responsible for group selection employ paradigms and population statistics borrowed from evolutionary game theory. Altruism is defined as "a concern for the welfare of others". If altruism is genetically determined, then altruistic individuals must reproduce their own altruistic genetic traits for altruism to survive, but when altruists lavish their resources on non-altruists at the expense of their own kind, the altruists tend to die out and the others tend to increase. An extreme example is a soldier losing his life trying to help a fellow soldier. This example raises the question of how altruistic genes can be passed on if this soldier dies without having any children.[10] Within sociobiology, a social behavior is first explained as a sociobiological hypothesis by finding an evolutionarily stable strategy that matches the observed behavior. Stability of a strategy can be difficult to prove, but usually, it will predict gene frequencies. The hypothesis can be supported by establishing a correlation between the gene frequencies predicted by the strategy, and those expressed in a population. Altruism between social insects and littermates has been explained in such a way. Altruistic behavior, behavior that increases the reproductive fitness of others at the apparent expense of the altruist,[11] in some animals has been correlated to the degree of genome shared between altruistic individuals. A quantitative description of infanticide by male harem-mating animals when the alpha male is displaced as well as rodent female infanticide and fetal resorption are active areas of study. In general, females with more bearing opportunities may value offspring less, and may also arrange bearing opportunities to maximize the food and protection from mates. An important concept in sociobiology is that temperament traits exist in an ecological balance. Just as an expansion of a sheep population might encourage the expansion of a wolf population, an expansion of altruistic traits within a gene pool may also encourage increasing numbers of individuals with dependent traits. Studies of human behavior genetics have generally found behavioral traits such as creativity, extroversion, aggressiveness, and IQ have high heritability. The researchers who carry out those studies are careful to point out that heritability does not constrain the influence that environmental or cultural factors may have on those traits.[12][13] Criminality is actively under study, but extremely controversial. There are arguments that in some environments criminal behavior might be adaptive.[14] The novelist Elias Canetti also has noted applications of sociobiological theory to cultural practices such as slavery and autocracy.[15]


Support for premise[edit] Genetic mouse mutants illustrate the power that genes exert on behaviour. For example, the transcription factor FEV (aka Pet1), through its role in maintaining the serotonergic system in the brain, is required for normal aggressive and anxiety-like behavior.[16] Thus, when FEV is genetically deleted from the mouse genome, male mice will instantly attack other males, whereas their wild-type counterparts take significantly longer to initiate violent behaviour. In addition, FEV has been shown to be required for correct maternal behaviour in mice, such that offspring of mothers without the FEV factor do not survive unless cross-fostered to other wild-type female mice.[17] A genetic basis for instinctive behavioural traits among non-human species, such as in the above example, is commonly accepted among many biologists; however, attempting to use a genetic basis to explain complex behaviours in human societies has remained extremely controversial.[18][19]


Reception[edit] Steven Pinker argues that critics have been overly swayed by politics and a fear of biological determinism,[a] accusing among others Stephen Jay Gould and Richard Lewontin of being "radical scientists", whose stance on human nature is influenced by politics rather than science,[21] while Lewontin, Steven Rose and Leon Kamin who drew a distinction between the politics and history of an idea and its scientific validity[22] argue that sociobiology fails on scientific grounds. Gould grouped sociobiology with eugenics, criticizing both in his book The Mismeasure of Man.[23] Noam Chomsky has expressed views on sociobiology on several occasions. During a 1976 meeting of the Sociobiology Study Group, as reported by Ullica Segerstråle, Chomsky argued for the importance of a sociobiologically informed notion of human nature.[24] Chomsky argued that human beings are biological organisms and ought to be studied as such, with his criticism of the "blank slate" doctrine in the social sciences (which would inspire a great deal of Steven Pinker's and others' work in evolutionary psychology), in his 1975 Reflections on Language.[25] Chomsky further hinted at the possible reconciliation of his anarchist political views and sociobiology in a discussion of Peter Kropotkin's Mutual Aid: A Factor of Evolution, which focused more on altruism than aggression, suggesting that anarchist societies were feasible because of an innate human tendency to cooperate.[26] Wilson has claimed that he had never meant to imply what ought to be, only what is the case. However, some critics have argued that the language of sociobiology readily slips from "is" to "ought",[22] an instance of the naturalistic fallacy. Pinker has argued that opposition to stances considered anti-social, such as ethnic nepotism, is based on moral assumptions, meaning that such opposition is not falsifiable by scientific advances.[27] The history of this debate, and others related to it, are covered in detail by Cronin (1993), Segerstråle (2000), and Alcock (2001).


See also[edit] Amity-Enmity Complex Biocultural anthropology Biosocial theory Cultural selection theory Darwinian anthropology Dual inheritance theory Ethics and evolutionary psychology Evolutionary anthropology Evolutionary developmental psychology Evolutionary ethics Evolutionary neuroscience Evolutionary psychology Genopolitics Human behavioral ecology Iterated prisoner's dilemma Kin selection Prisoner's dilemma Social evolution Social neuroscience Sociophysiology


Notes[edit] ^ Biological determinism was a right wing philosophy underlying the social Darwinian and eugenics movements of the early 20th century, and controversies in the history of intelligence testing.[20]


References[edit] ^ Wilson, E.O. (1978). On Human Nature, Page x, Cambridge, Ma: Harvard ^ David Sloan Wilson and Edward O. Wilson (2007), "Rethinking The Theoretical Foundation of Sociobiology", The Quarterly Review of Biology, 82 (4): 327–348, doi:10.1086/522809  ^ Packer, Craig; Pusey, Anne E. (1983). "Adaptations of Female Lions to Infanticide by Incoming Males" (PDF). Am. Nat. 121: 716–728.  ^ "The Life of J.P. Scott". Bowling Green State University. Retrieved 14 December 2016.  ^ Midgley, Mary (September 1984), "Sociobiology", Journal of Medical Ethics, BMJ Publishing Group, 10 (3): 158–160, doi:10.1136/jme.10.3.158, JSTOR 27716288  ^ Conniff, Richard (June 2006). "Discover Interview: E.O. Wilson". Discover Magazine. Retrieved 21 December 2011.  ^ Walsh, Bryan (17 August 2011). "All-TIME 100 Nonfiction Books".  ^ The Handbook of Evolutionary Psychology, edited by David M. Buss, John Wiley & Sons, Inc., 2005. Chapter 5 by Edward H. Hagen . ^ Wilson, 1975. Chapter 5. "Group Selection and Altruism" ^ Tessman, Irwin (1995). "Human altruism as a courtship display". Forum: 157.  ^ Costa-Leonarda, Ana Maria (2014). "Termite Communication during different behavioral activities". in: Witzany G (ed). Biocommunication of Animals: 161–190.  ^ Johnson, Wendy; Turkheimer, E.; Gottesman, Irving; Bouchard, Thomas (2009). "Beyond Heritability: Twin Studies in Behavioral Research" (PDF). Current Directions in Psychological Science. 18 (4): 217–220. doi:10.1111/j.1467-8721.2009.01639.x. PMC 2899491 . PMID 20625474. Retrieved 29 June 2010. Moreover, even highly heritable traits can be strongly manipulated by the environment, so heritability has little if anything to do with controllability. For example, height is on the order of 90% heritable, yet North and South Koreans, who come from the same genetic background, presently differ in average height by a full 6 inches (Pak, 2004; Schwekendiek, 2008).  ^ Turkheimer, Eric (April 2008). "A Better Way to Use Twins for Developmental Research" (PDF). LIFE Newsletter. Max Planck Institute for Human Development. 2 (1): 2–5. Retrieved 29 October 2010. But back to the question: What does heritability mean? Almost everyone who has ever thought about heritability has reached a commonsense intuition about it: One way or another, heritability has to be some kind of index of how genetic a trait is. That intuition explains why so many thousands of heritability coefficients have been calculated over the years. . . . Unfortunately, that fundamental intuition is wrong. Heritability isn't an index of how genetic a trait is. A great deal of time has been wasted in the effort of measuring the heritability of traits in the false expectation that somehow the genetic nature of psychological phenomena would be revealed.  ^ The Sociobiology Of Sociopathy: An Integrated ^ Elias Canetti, Crowds and Power. Harmondsworth: Penguin, 1981, p. 444-445. ^ Hendricks TJ, Fyodorov DV, Wegman LJ, Lelutiu NB, Pehek EA, Yamamoto B, Silver J, Weeber EJ, Sweatt JD, Deneris ES. Pet-1 ETS gene plays a critical role in 5-HT neuron development and is required for normal anxiety-like and aggressive behaviour]. Neuron. 2003 Jan 23;37(2):233-47 ^ Lerch-Haner JK, Frierson D, Crawford LK, Beck SG, Deneris ES. Serotonergic transcriptional programming determines maternal behavior and offspring survival. Nat Neurosci. 2008 Sep;11(9):1001-3. ^ Fisher, Helen (16 October 1994). "'Wilson,' They Said, 'Your All Wet!'". New York Times. Retrieved 21 July 2015.  ^ Gould, Stephen Jay (16 November 1978). "Sociobiology: the art of storytelling". New Scientist. 80 (1129): 530–533.  ^ Allen, Garland E. (1984). "The Roots of Biological Determinism: review of The Mismeasure of Man by Stephen Jay Gould". Journal of the History of Biology. 17 (1): 141–145. JSTOR 4330882.  ^ Pinker, Steven (2002). The Blank Slate: The Modern Denial of Human Nature. New York: Penguin Books. p. 149. ISBN 0-14-200334-4. A surprising number of intellectuals, particularly on the left, do deny that there is such a thing as inborn talent, especially intelligence. Stephen Jay Gould's 191 bestseller The Mismeasure of Man was written to debunk 'the abstraction of intelligence as a single entity ... and the use of these numbers to rank people in a single series of worthiness'  ^ a b Richard Lewontin; Leon Kamin; Steven Rose (1984). Not in Our Genes: Biology, Ideology, and Human Nature. Pantheon Books. ISBN 0-394-50817-3.  ^ Gould, Stephen Jay (1996). The Mismeasure of Man. p. Introduction to the Revised Edition.  ^ Segerstråle 2000, p. 205. ^ Chomsky, Noam (1975), Reflections on Language:10. New York: Pantheon Books. ^ Chomsky (1995). "Rollback, Part II." Z Magazine 8 (Feb.): 20–31. ^ Pinker, Steven (2002). The Blank Slate: The Modern Denial of Human Nature. New York: Viking. p. 145 Bibliography[edit] Alcock, John (2001). The triumph of sociobiology. Oxford, UK: Oxford University Press. ISBN 978-0-19-514383-6.  Barkow, Jerome (Ed.). (2006) Missing the Revolution: Darwinism for Social Scientists. Oxford: Oxford University Press. Cronin, Helena (1993). The ant and the peacock: Altruism and sexual selection from Darwin to today. Press Syndicate of the University of Cambridge. ISBN 978-0-521-45765-1.  Nancy Etcoff (1999). Survival of the Prettiest: The Science of Beauty. Anchor Books. ISBN 0-385-47942-5.  Haugan, Gørill (2006) Nursing home patients’ spirituality. Interaction of the spiritual, physical, emotional and social dimensions (Faculty of Nursing, Sør-Trøndelag University College Norwegian University of Science and Technology) Richard M. Lerner (1992). Final Solutions: Biology, Prejudice, and Genocide. Pennsylvania State University Press. ISBN 0-271-00793-1.  Richards, Janet Radcliffe (2000). Human Nature After Darwin: A Philosophical Introduction. London: Routledge. Segerstråle, Ullica (2000). Defenders of the truth: The sociobiology debate. Oxford, UK: Oxford University Press. ISBN 978-0-19-286215-0.  Gisela Kaplan; Lesley J Rogers (2003). Gene Worship: Moving Beyond the Nature/Nurture Debate over Genes, Brain, and Gender. Other Press. ISBN 1-59051-034-8.  F. H. Schmidt: Verhaltensforschung und Recht, Berlin, Duncker & Humblot, 1982, ISBN 3 428 05099 1


External links[edit] Wikimedia Commons has media related to Sociobiology. Sociobiology (Stanford Encyclopedia of Philosophy) - Harmon Holcomb & Jason Byron The Sociobiology of Sociopathy, Mealey, 1995 Speak, Darwinists! Interviews with leading sociobiologists. Race and Creation - Richard Dawkins Genetic Similarity and Ethnic Nationalism - An Attempted Sociobiological Explanation of the scientific basis for Political Group Formation. Scientist at Work | Edward O. Wilson | Taking a Cue From Ants on Evolution of Humans by Nicholas Wade v t e Sociobiology Topics Behavioural genetics Challenge hypothesis Dual inheritance theory Ethology Evolutionary psychology Evolution of morality Evolutionary models of food sharing Group selection Kin recognition Kin selection Male Warrior hypothesis Reciprocal altruism Sexual selection in human evolution Sex and psychology Sociality (Eusociality, Evolution of eusociality, Presociality) Dunbar's number Supporters Anne Campbell Noam Chomsky Richard Dawkins Daniel Dennett Sarah Blaffer Hrdy Steven Pinker Frans de Waal E. O. Wilson Sociobiology: The New Synthesis Robert Wright Opponents Stephen Jay Gould Leon Kamin Richard Lewontin Steven Rose Not in Our Genes Evolutionary biology portal v t e Evolutionary psychology Processes Altruism Group selection Kin selection Sexual selection Sociobiology Coevolution Evolutionarily stable strategy Areas Psychological development Morality Religion Depression Developmental psychopathology Educational psychology Evolutionary aesthetics Music Darwinian literary studies Evolution of emotion Biologists/ Neuroscientists John Crook Charles Darwin Richard Dawkins Jared Diamond W. D. Hamilton Peter Kropotkin Gordon Orians Jaak Panksepp Margie Profet Giacomo Rizzolatti Randy Thornhill Robert Trivers Carel van Schaik Claus Wedekind Wolfgang Wickler David Sloan Wilson E. O. Wilson George C. Williams Anthropologists Jerome H. Barkow Robert Boyd Robin Dunbar Daniel Fessler Mark Flinn Joseph Henrich Ruth Mace Daniel Nettle Frank Salter Stephen Shennan Donald Symons John Tooby Behavioral economists/ Political scientists Samuel Bowles Ernst Fehr Herbert Gintis Dominic D. P. Johnson Gad Saad Literary theory/ Aesthetics Joseph Carroll Denis Dutton Psychologists/ Cognitive scientists Simon Baron-Cohen Justin L. Barrett Jay Belsky David F. Bjorklund Paul Bloom Pascal Boyer Joseph Bulbulia David Buss Josep Call Anne Campbell Leda Cosmides Martin Daly Daniel Dennett Paul Ekman Anne Fernald David C. Geary Gerd Gigerenzer Jonathan Haidt Judith Rich Harris Aurelio José Figueredo Stephen Kaplan Douglas T. Kenrick Simon M. Kirby Robert Kurzban Michael T. McGuire Geoffrey Miller Randolph M. Nesse Steven Neuberg David Perrett Steven Pinker Paul Rozin Mark Schaller David P. Schmitt Todd K. Shackelford Roger Shepard Peter K. Smith Dan Sperber Anthony Stevens Frank Sulloway Michael Tomasello Mark van Vugt Glenn Wilson Margo Wilson Related subjects and articles The Adapted Mind The Evolution of Human Sexuality Evolutionary Psychology Evolution and Human Behavior Memetics Sociobiology Evolutionary neuroscience Human evolution Sociocultural evolution Evolutionary anthropology Evolutionary medicine Evolutionary linguistics Evolutionary psychology and culture Biosocial criminology Criticism of evolutionary psychology Lists Evolutionary psychologists Evolutionary psychology research groups and centers Bibliography of evolution and human behavior Evolutionary psychology Psychology portal Evolutionary biology portal v t e Ethology Branches Animal cognition Animal communication Animal consciousness Animal culture Animal sexual behaviour Animal welfare science Anthrozoology Bee learning and communication Behavioural ecology Behavioural genetics Cognitive ethology Comparative psychology Emotion in animals Evolutionary neuroscience Human ethology Instinct Learning Neuroethology Pain in animals Sociobiology Tool use by animals Zoosemiotics Zoomusicology Ethologists Patrick Bateson Marc Bekoff Donald Broom Charles Darwin Marian Dawkins Richard Dawkins Irenäus Eibl-Eibesfeldt Dian Fossey Karl von Frisch Jane Goodall Heini Hediger Julian Huxley Konrad Lorenz Desmond Morris Thomas Sebeok William Homan Thorpe Nikolaas Tinbergen Jakob von Uexküll Wolfgang Wickler E. O. Wilson Solly Zuckerman Societies Association for the Study of Animal Behaviour International Society for Applied Ethology Journals Animal Behaviour Animal Cognition Animal Welfare Behavioral Ecology Behaviour Related topics Animals portal Biology portal  Ethology (category) v t e Nikolaas Tinbergen Selected works The Study of Instinct (book) The Herring Gull's World (book) Social Behaviour in Animals: With Special Reference to Vertebrates (book) Signals for Survival (film) The Riddle of the Rook (film) General Animal behaviour science Behavioural biology Behavioral ecology History of ethology Neuroethology Sociobiology Hawk/goose effect Supernormal stimulus Tinbergen's four questions Related Jan Tinbergen Luuk Tinbergen Konrad Lorenz Karl von Frisch Richard Dawkins Desmond Morris 1973 Nobel Prize in Physiology or Medicine Authority control NDL: 00575496 Retrieved from "https://en.wikipedia.org/w/index.php?title=Sociobiology&oldid=820588299" Categories: SociobiologyBiologyEcologyEthologyEvolutionary biologyEvolutionary psychologySubfields of zoologyScience studiesHidden categories: Pages using div col without cols and colwidth parameters


Navigation menu Personal tools Not logged inTalkContributionsCreate accountLog in Namespaces ArticleTalk Variants Views ReadEditView history More Search Navigation Main pageContentsFeatured contentCurrent eventsRandom articleDonate to WikipediaWikipedia store Interaction HelpAbout WikipediaCommunity portalRecent changesContact page Tools What links hereRelated changesUpload fileSpecial pagesPermanent linkPage informationWikidata itemCite this page Print/export Create a bookDownload as PDFPrintable version In other projects Wikimedia CommonsWikiquote Languages العربيةБеларускаяCatalàČeštinaDanskDeutschEestiEspañolفارسیFrançaisGalego한국어IdoBahasa IndonesiaItalianoעבריתҚазақшаLietuviųMagyarBahasa MelayuNederlands日本語NorskPiemontèisPolskiPortuguêsRomânăРусскийSimple EnglishSlovenčinaSlovenščinaСрпски / srpskiSrpskohrvatski / српскохрватскиSuomiSvenskaTürkçeУкраїнськаاردوWinaray中文 Edit links This page was last edited on 15 January 2018, at 14:01. Text is available under the Creative Commons Attribution-ShareAlike License; additional terms may apply. By using this site, you agree to the Terms of Use and Privacy Policy. Wikipedia® is a registered trademark of the Wikimedia Foundation, Inc., a non-profit organization. Privacy policy About Wikipedia Disclaimers Contact Wikipedia Developers Cookie statement Mobile view (window.RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgPageParseReport":{"limitreport":{"cputime":"0.396","walltime":"0.504","ppvisitednodes":{"value":2341,"limit":1000000},"ppgeneratednodes":{"value":0,"limit":1500000},"postexpandincludesize":{"value":116586,"limit":2097152},"templateargumentsize":{"value":1737,"limit":2097152},"expansiondepth":{"value":15,"limit":40},"expensivefunctioncount":{"value":0,"limit":500},"entityaccesscount":{"value":1,"limit":400},"timingprofile":["100.00% 378.251 1 -total"," 40.18% 151.993 2 Template:Reflist"," 11.34% 42.893 7 Template:Cite_journal"," 10.47% 39.610 2 Template:Citation"," 9.87% 37.339 9 Template:Cite_book"," 8.75% 33.103 4 Template:Navbox"," 7.70% 29.116 1 Template:Evolutionary_biology"," 7.41% 28.045 1 Template:Commonscat"," 7.14% 27.001 1 Template:Sidebar_with_collapsible_lists"," 6.10% 23.057 1 Template:Commons"]},"scribunto":{"limitreport-timeusage":{"value":"0.164","limit":"10.000"},"limitreport-memusage":{"value":4837764,"limit":52428800}},"cachereport":{"origin":"mw1261","timestamp":"20180115175152","ttl":1900800,"transientcontent":false}}});});(window.RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgBackendResponseTime":73,"wgHostname":"mw1244"});});


Sociobiological - Photos and All Basic Informations

Sociobiological More Links

Sociobiology: The New SynthesisEvolutionGenetic DivergenceTaxonomy (biology)Common DescentIntroduction To EvolutionCommon DescentEvidence Of Common DescentPopulation GeneticsGenetic VariationGenetic DiversityMutationNatural SelectionAdaptationPolymorphism (biology)Genetic DriftGene FlowSpeciationAdaptive RadiationCo-operation (evolution)CoevolutionDivergent EvolutionConvergent EvolutionParallel EvolutionExtinctionAbiogenesisEvolutionary History Of LifeTimeline Of Evolutionary History Of LifeHuman EvolutionPhylogenetic TreeBiodiversityBiogeographyTaxonomy (biology)Evolutionary TaxonomyCladisticsTransitional FossilExtinction EventHistory Of Evolutionary ThoughtEvolutionary Ideas Of The Renaissance And EnlightenmentTransmutation Of SpeciesCharles DarwinOn The Origin Of SpeciesThe Eclipse Of DarwinismModern Synthesis (20th Century)History Of Molecular EvolutionEvolutionary Developmental BiologyEvolutionary BiologyHistory Of PaleontologyTimeline Of PaleontologyApplications Of EvolutionBiosocial CriminologyEcological GeneticsEvolutionary AestheticsEvolutionary AnthropologyEvolutionary ComputationEvolutionary EcologyEvolutionary EconomicsEvolutionary EpistemologyEvolutionary EthicsEvolutionary Game TheoryEvolutionary LinguisticsEvolutionary MedicineEvolutionary NeuroscienceEvolutionary PhysiologyEvolutionary PsychologyExperimental EvolutionPhylogeneticsPaleontologySelective BreedingSystematicsUniversal DarwinismEvolution As Fact And TheorySocial Effects Of Evolutionary TheoryCreation–evolution ControversyObjections To EvolutionLevel Of Support For EvolutionPortal:Evolutionary BiologyCategory:Evolutionary BiologyBook:EvolutionIndex Of Evolutionary Biology ArticlesTemplate:Evolutionary BiologyTemplate Talk:Evolutionary BiologyBiologySocial BehaviorEvolutionEthologyAnthropologyEvolutionZoologyArchaeologyPopulation GeneticsSocietyDarwinian AnthropologyHuman Behavioral EcologyEvolutionary PsychologyHive (disambiguation)Social InsectE. O. WilsonSociobiology: The New SynthesisScientific ControversyRichard LewontinStephen Jay GouldNature And NurtureNatural SelectionHeredityAdaptive BehaviourCompetition (biology)EnlargeE. O. WilsonSociobiology: The New SynthesisJohn Paul Scott (geneticist)Peter KropotkinMutual Aid: A Factor Of EvolutionRichard D. AlexanderRobert TriversWilliam D. HamiltonJ. B. S. HaldaneAltruismAltruismAggressionHymenopteraPulitzer PrizeOn Human NatureNature (journal)Science (journal)Behavioral EcologyHuman NatureNatural SelectionOrganismSpeciesEvolutionary BiologyEnlargeNikolaas TinbergenNikolaas TinbergenTinbergen's Four QuestionsAdaptationPhylogenyOntogenyBrain AnatomyHormoneInstinctIntuition (knowledge)MammalsOffspringAdaptationSocial GroupGroup SelectionParadigmsEvolutionary Game TheoryHypothesisEvolutionarily Stable StrategySocial InsectGenomeInfanticideAlpha MaleRodentFoodEcologicalSheepWolfIntelligence QuotientHeritabilityCrimeElias CanettiTranscription FactorSerotonergic SystemAggressionAnxietySteven PinkerBiological DeterminismStephen Jay GouldRichard LewontinSteven RoseLeon KaminEugenicsThe Mismeasure Of ManNoam ChomskySociobiology Study GroupBlank SlatePeter KropotkinMutual Aid: A Factor Of EvolutionNaturalistic FallacyEthnic NepotismMoralityFalsifiableAmity-Enmity ComplexBiocultural AnthropologyBiosocial TheoryCultural Selection TheoryDarwinian AnthropologyDual Inheritance TheoryEthics And Evolutionary PsychologyEvolutionary AnthropologyEvolutionary Developmental PsychologyEvolutionary EthicsEvolutionary NeuroscienceEvolutionary PsychologyGenopoliticsHuman Behavioral EcologyIterated Prisoner's DilemmaKin SelectionPrisoner's DilemmaSocial EvolutionSocial NeuroscienceSociophysiologySocial DarwinismEugenicsHistory Of The Race And Intelligence ControversyDigital Object IdentifierDigital Object IdentifierJSTORDigital Object IdentifierPubMed CentralPubMed IdentifierElias CanettiCrowds And PowerJSTORSteven PinkerThe Blank SlateInternational Standard Book NumberSpecial:BookSources/0-14-200334-4International Standard Book NumberSpecial:BookSources/0-394-50817-3Stephen Jay GouldInternational Standard Book NumberSpecial:BookSources/978-0-19-514383-6International Standard Book NumberSpecial:BookSources/978-0-521-45765-1Nancy EtcoffInternational Standard Book NumberSpecial:BookSources/0-385-47942-5Richard M. LernerInternational Standard Book NumberSpecial:BookSources/0-271-00793-1International Standard Book NumberSpecial:BookSources/978-0-19-286215-0International Standard Book NumberSpecial:BookSources/1-59051-034-8International Standard Book NumberSpecial:BookSources/3+428+05099+1Richard DawkinsTemplate:SociobiologyTemplate Talk:SociobiologyBehavioural GeneticsChallenge HypothesisDual Inheritance TheoryEthologyEvolutionary PsychologyEvolution Of MoralityEvolutionary Models Of Food SharingGroup SelectionKin RecognitionKin SelectionMale Warrior HypothesisReciprocal AltruismSexual Selection In Human EvolutionSex And PsychologySocialityEusocialityEvolution Of EusocialityPresocialityDunbar's NumberAnne Campbell (academic)Noam ChomskyRichard DawkinsDaniel DennettSarah Blaffer HrdySteven PinkerFrans De WaalE. O. WilsonSociobiology: The New SynthesisRobert Wright (journalist)Stephen Jay GouldLeon KaminRichard LewontinSteven RoseNot In Our GenesPortal:Evolutionary BiologyTemplate:Evolutionary PsychologyTemplate Talk:Evolutionary PsychologyEvolutionary PsychologyReciprocal AltruismGroup SelectionKin SelectionSexual Selection In HumansCoevolutionEvolutionarily Stable StrategyEvolutionary Developmental PsychologyEvolution Of MoralityEvolutionary Psychology Of ReligionEvolutionary Approaches To DepressionEvolutionary Developmental PsychopathologyEvolutionary Educational PsychologyEvolutionary AestheticsEvolutionary MusicologyDarwinian Literary StudiesEvolution Of EmotionBiologyNeuroscienceJohn Crook (ethologist)Charles DarwinRichard DawkinsJared DiamondW. D. HamiltonPeter KropotkinGordon OriansJaak PankseppMargie ProfetGiacomo RizzolattiRandy ThornhillRobert TriversCarel Van SchaikClaus WedekindWolfgang WicklerDavid Sloan WilsonE. O. WilsonGeorge C. Williams (biologist)AnthropologyJerome H. BarkowRobert Boyd (anthropologist)Robin DunbarDaniel FesslerMark FlinnJoseph HenrichRuth MaceDaniel NettleFrank SalterStephen ShennanDonald SymonsJohn ToobyBehavioral EconomicsPolitical ScienceSamuel Bowles (economist)Ernst FehrHerbert GintisDominic D. P. JohnsonGad SaadLiterary TheoryAestheticsJoseph Carroll (scholar)Denis DuttonPsychologyCognitive ScienceSimon Baron-CohenJustin L. BarrettJay BelskyDavid F. BjorklundPaul Bloom (psychologist)Pascal BoyerJoseph BulbuliaDavid BussJosep CallAnne Campbell (academic)Leda CosmidesMartin DalyDaniel DennettPaul EkmanAnne FernaldDavid C. GearyGerd GigerenzerJonathan HaidtJudith Rich HarrisAurelio José FigueredoRachel And Stephen KaplanDouglas T. KenrickSimon M. KirbyRobert KurzbanMichael T. McGuireGeoffrey Miller (psychologist)Randolph M. NesseSteven NeubergDavid PerrettSteven PinkerPaul RozinMark SchallerDavid P. SchmittTodd K. ShackelfordRoger ShepardPeter K. SmithDan SperberAnthony Stevens (Jungian Analyst)Frank SullowayMichael TomaselloMark Van VugtGlenn Wilson (psychologist)Margo WilsonThe Adapted MindThe Evolution Of Human SexualityEvolutionary Psychology (journal)Evolution And Human BehaviorMemeticsEvolutionary NeuroscienceHuman EvolutionSociocultural EvolutionEvolutionary AnthropologyEvolutionary MedicineEvolutionary LinguisticsEvolutionary Psychology And CultureBiosocial CriminologyCriticism Of Evolutionary PsychologyList Of Evolutionary PsychologistsEvolutionary Psychology Research Groups And CentersEvolutionary PsychologyCategory:Evolutionary PsychologyPortal:PsychologyPortal:Evolutionary BiologyTemplate:EthologyTemplate Talk:EthologyEthologyAnimal CognitionAnimal CommunicationAnimal ConsciousnessAnimal CultureAnimal Sexual BehaviourAnimal Welfare ScienceAnthrozoologyBee Learning And CommunicationBehavioral EcologyBehavioural GeneticsCognitive EthologyComparative PsychologyEmotion In AnimalsEvolutionary NeuroscienceHuman EthologyInstinctLearningNeuroethologyPain In AnimalsTool Use By AnimalsZoosemioticsZoomusicologyPatrick BatesonMarc BekoffDonald BroomCharles DarwinMarian DawkinsRichard DawkinsIrenäus Eibl-EibesfeldtDian FosseyKarl Von FrischJane GoodallHeini HedigerJulian HuxleyKonrad LorenzDesmond MorrisThomas SebeokWilliam Homan ThorpeNikolaas TinbergenJakob Von UexküllWolfgang WicklerE. O. WilsonSolly ZuckermanAssociation For The Study Of Animal BehaviourInternational Society For Applied EthologyAnimal Behaviour (journal)Animal CognitionAnimal Welfare (journal)Behavioral Ecology (journal)Behaviour (journal)Portal:AnimalsPortal:BiologyCategory:EthologyTemplate:Nikolaas TinbergenNikolaas TinbergenThe Study Of InstinctAnimal BehaviourBehavioural BiologyBehavioral EcologyHistory Of EthologyNeuroethologyHawk/goose EffectSupernormal StimulusTinbergen's Four QuestionsJan TinbergenLuuk TinbergenKonrad LorenzKarl Von FrischRichard DawkinsDesmond MorrisNobel Prize In Physiology Or MedicineHelp:Authority ControlNational Diet LibraryHelp:CategoryCategory:SociobiologyCategory:BiologyCategory:EcologyCategory:EthologyCategory:Evolutionary BiologyCategory:Evolutionary PsychologyCategory:Subfields Of ZoologyCategory:Science StudiesCategory:Pages Using Div Col Without Cols And Colwidth ParametersDiscussion About Edits From This IP Address [n]A List Of Edits Made From This IP Address [y]View The Content Page [c]Discussion About The Content Page [t]Edit This Page [e]Visit The Main Page [z]Guides To Browsing WikipediaFeatured Content – The Best Of WikipediaFind Background Information On Current EventsLoad A Random Article [x]Guidance On How To Use And Edit WikipediaFind Out About WikipediaAbout The Project, What You Can Do, Where To Find ThingsA List Of Recent Changes In The Wiki [r]List Of All English Wikipedia Pages Containing Links To This Page [j]Recent Changes In Pages Linked From This Page [k]Upload Files [u]A List Of All Special Pages [q]Wikipedia:AboutWikipedia:General Disclaimer



view link view link view link view link view link