Contents 1 Description 2 Etymology 3 See also 4 References 5 External links


Description[edit] This section needs expansion. You can help by adding to it. (June 2015) Harvested rhubarb petioles with leaves attached The petiole is a stalk that attaches a leaf to the plant stem. In petiolate leaves, the leaf stalk (petiole) may be long, as in the leaves of celery and rhubarb, short or completely absent, in which case the blade attaches directly to the stem and is said to be sessile. Subpetiolate leaves are nearly petiolate, or have an extremely short petiole, and may appear sessile.[2]:157 The broomrape family Orobanchaceae is an example of a family in which the leaves are always sessile.[3]:639 In some other plant groups, such as the speedwell genus Veronica, petiolate and sessile leaves may occur in different species.[3]:584 In the grasses (Poaceae) the leaves are apetiolate, but the leaf blade may be narrowed at the junction with the leaf sheath to form a pseudopetiole, as in Pseudosasa japonica.[4]:391 In plants with compound leaves, the leaflets are attached to a continuation of the petiole called the rachis.[1]:98 Each leaflet may be attached to the rachis by a short stalk called the petiolule.[1]:87 There may be swollen regions at either end of the petiole known as pulvina (singular=pulvinus)[1]:97 that are composed of a flexible tissue that allows leaf movement. Pulvina are common in the bean family Fabaceae and the prayer plant family Marantaceae. A pulvinus on a petiolule is called a pulvinulus. In some plants, the petioles are flattened and widened, to become phyllodes or phyllodia, or cladophylls and the true leaves may be reduced or absent. Thus, the phyllode comes to serve the functions of the leaf. Phyllodes are common in the genus Acacia, especially the Australian species, at one time put in Acacia subgenus Phyllodineae. Acacia koa with phyllode between the branch and the compound leaves In Acacia koa, the phyllodes are leathery and thick, allowing the tree to survive stressful environments. The petiole allows partially submerged hydrophytes to have leaves floating at different depths, the petiole being between the node and the stem. In plants such as rhubarb (Rheum rhabarbarum), celery (Apium graveolens), artichokes and cardoons (Cynara cardunculus) the petioles ("stalks" or "ribs") are cultivated as edible crops. The petiole of rhubarb grows directly from the rhizome and produces the leaf at its end. Botanically it is categorized as a vegetable and culinarily used as a fruit.[5][6]


Etymology[edit] Petiole is pronounced /ˈpiːtɪoʊl/ and comes from Latin petiolus, or peciolus "little foot", "stem", an alternative diminutive of pes "foot". The regular diminutive pediculus is also used for "foot stalk".


See also[edit] Hyponastic response


References[edit] ^ a b c d Beentje, H. The Kew plant glossary. London: Kew Publishing. ISBN 9781842464229.  ^ a b Mauseth, James D (2003). Botany: An Introduction to Plant Biology. Jones & Bartlett Learning. ISBN 0-7637-2134-4.  ^ a b Stace, C. A. (2010). New Flora of the British Isles (Third ed.). Cambridge, U.K.: Cambridge University Press. ISBN 9780521707725.  ^ Heywood, V.H.; Brummitt, R.K.; Culham, A.; Seberg, O. (2007). Flowering plant families of the world. New York: Firefly Books. ISBN 9781554072064.  ^ Foust, Clifford M. (1992). Rhubarb: The Wondrous Drug. Princeton: Princeton University Press. ISBN 0-691-08747-4.  ^ High Altitude Rhubarb


External links[edit]  "Petiole". Collier's New Encyclopedia. 1921.  Wikimedia Commons has media related to petioles. v t e Botany History of botany Subdisciplines Plant systematics Ethnobotany Paleobotany Plant anatomy Plant ecology Phytogeography Geobotany Flora Phytochemistry Plant pathology Bryology Phycology Floristics Dendrology Plant groups Algae Archaeplastida Bryophyte Non-vascular plants Vascular plants Spermatophytes Pteridophyte Gymnosperm Angiosperm Plant morphology (glossary) Plant cells Cell wall Phragmoplast Plastid Plasmodesma Vacuole Tissues Meristem Vascular tissue Vascular bundle Ground tissue Mesophyll Cork Wood Storage organs Vegetative Root Rhizoid Bulb Rhizome Shoot Stem Leaf Petiole Cataphyll Bud Sessility Reproductive (Flower) Flower development Inflorescence Umbel Raceme Bract Pedicellate Flower Whorl Floral symmetry Floral diagram Floral formula Receptacle Hypanthium (Floral cup) Perianth Tepal Petal Sepal Sporophyll Gynoecium Ovary Ovule Stigma Archegonium Androecium Stamen Staminode Pollen Tapetum Gynandrium Gametophyte Sporophyte Plant embryo Fruit Fruit anatomy Berry Capsule Seed Seed dispersal Endosperm Surface structures Epicuticular wax Plant cuticle Epidermis Stoma Nectary Trichome Prickle Plant physiology Materials Nutrition Photosynthesis Chlorophyll Plant hormone Transpiration Turgor pressure Bulk flow Aleurone Phytomelanin Sugar Sap Starch Cellulose Plant growth and habit Secondary growth Woody plants Herbaceous plants Habit Vines Lianas Shrubs Subshrubs Trees Succulent plants Reproduction Evolution Ecology Alternation of generations Sporangium Spore Microsporangia Microspore Megasporangium Megaspore Pollination Pollinators Pollen tube Double fertilization Germination Evolutionary development Evolutionary history timeline Hardiness zone Plant taxonomy History of plant systematics Herbarium Biological classification Botanical nomenclature Botanical name Correct name Author citation International Code of Nomenclature for algae, fungi, and plants (ICN) - for Cultivated Plants (ICNCP) Taxonomic rank International Association for Plant Taxonomy (IAPT) Plant taxonomy systems Cultivated plant taxonomy Citrus taxonomy cultigen cultivar Group grex Practice Agronomy Floriculture Forestry Horticulture Lists Related topics Botanical terms Botanists by author abbreviation Botanical expedition Category Portal WikiProject Retrieved from "https://en.wikipedia.org/w/index.php?title=Petiole_(botany)&oldid=816324066" Categories: Plant morphologyHidden categories: Articles to be expanded from June 2015All articles to be expandedArticles using small message boxesWikipedia articles incorporating a citation from Collier's EncyclopediaCommons category with local link different than on Wikidata


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