Contents 1 Etymology 2 Definition 3 Diversity 4 Hypothetical ancestral mollusc 4.1 Mantle and mantle cavity 4.2 Shell 4.3 Foot 4.4 Circulatory system 4.5 Respiration 4.6 Eating, digestion, and excretion 4.7 Nervous system 4.8 Reproduction 5 Ecology 5.1 Feeding 6 Classification 7 Evolution 7.1 Fossil record 7.2 Phylogeny 8 Human interaction 8.1 Uses by humans 8.1.1 Bioindicators 8.2 Harmful to humans 8.2.1 Stings and bites 8.2.2 Disease vectors 8.2.3 Pests 9 Notes 10 References 11 Bibliography 12 Further reading 13 External links

Etymology[edit] The words mollusc and mollusk are both derived from the French mollusque, which originated from the Latin molluscus, from mollis, soft. Molluscus was itself an adaptation of Aristotle's τα μαλακά (ta malaká), "the soft things", which he applied to cuttlefish. The scientific study of molluscs is accordingly called malacology.[4] The name Molluscoida was formerly used to denote a division of the animal kingdom containing the brachiopods, bryozoans, and tunicates, the members of the three groups having been supposed to somewhat resemble the molluscs. As it is now known these groups have no relation to molluscs, and very little to one another, the name Molluscoida has been abandoned.[5]

Definition[edit] The most universal features of the body structure of molluscs are a mantle with a significant cavity used for breathing and excretion, and the organization of the nervous system. Many have a calcareous shell.[6] Molluscs have developed such a varied range of body structures, it is difficult to find synapomorphies (defining characteristics) to apply to all modern groups.[7] The most general characteristic of molluscs is they are unsegmented and bilaterally symmetrical.[8] The following are present in all modern molluscs:[9][10] The dorsal part of the body wall is a mantle (or pallium) which secretes calcareous spicules, plates or shells. It overlaps the body with enough spare room to form a mantle cavity. The anus and genitals open into the mantle cavity. There are two pairs of main nerve cords.[10] Other characteristics that commonly appear in textbooks have significant exceptions:   Whether characteristic is found in these classes of Molluscs Supposed universal Molluscan characteristic[9] Aplacophora[11] Polyplacophora[12] Monoplacophora[13] Gastropoda[14] Cephalopoda[15] Bivalvia[16] Scaphopoda[17] Radula, a rasping "tongue" with chitinous teeth Absent in 20% of Neomeniomorpha Yes Yes Yes Yes No Internal, cannot extend beyond body Broad, muscular foot Reduced or absent Yes Yes Yes Modified into arms Yes Small, only at "front" end Dorsal concentration of internal organs (visceral mass) Not obvious Yes Yes Yes Yes Yes Yes Large digestive ceca No ceca in some Aplacophora Yes Yes Yes Yes Yes No Large complex metanephridia ("kidneys") None Yes Yes Yes Yes Yes Small, simple One or more valves/ shells Primitive forms, yes; modern forms, no Yes Yes Snails, yes; slugs, mostly yes (internal vestigial) Octopuses, no; cuttlefish, nautilus, squid, yes Yes Yes Odontophore Yes Yes Yes Yes Yes No Yes

Diversity[edit] About 80% of all known mollusc species are gastropods (snails and slugs), including this cowry (a sea snail).[18] Estimates of accepted described living species of molluscs vary from 50,000 to a maximum of 120,000 species.[1] In 1969 David Nicol estimated the probable total number of living mollusc species at 107,000 of which were about 12,000 fresh-water gastropods and 35,000 terrestrial. The Bivalvia would comprise about 14% of the total and the other five classes less than 2% of the living molluscs.[19] In 2009, Chapman estimated the number of described living species at 85,000.[1] Haszprunar in 2001 estimated about 93,000 named species,[20] which include 23% of all named marine organisms.[21] Molluscs are second only to arthropods in numbers of living animal species[18]—far behind the arthropods' 1,113,000 but well ahead of chordates' 52,000.[22] About 200,000 living species in total are estimated,[1][23] and 70,000 fossil species,[9] although the total number of mollusc species ever to have existed, whether or not preserved, must be many times greater than the number alive today.[24] Molluscs have more varied forms than any other animal phylum. They include snails, slugs and other gastropods; clams and other bivalves; squids and other cephalopods; and other lesser-known but similarly distinctive subgroups. The majority of species still live in the oceans, from the seashores to the abyssal zone, but some form a significant part of the freshwater fauna and the terrestrial ecosystems. Molluscs are extremely diverse in tropical and temperate regions, but can be found at all latitudes.[7] About 80% of all known mollusc species are gastropods.[18] Cephalopoda such as squid, cuttlefish, and octopuses are among the neurologically most advanced of all invertebrates.[25] The giant squid, which until recently had not been observed alive in its adult form,[26] is one of the largest invertebrates, but a recently caught specimen of the colossal squid, 10 m (33 ft) long and weighing 500 kg (1,100 lb), may have overtaken it.[27] Freshwater and terrestrial molluscs appear exceptionally vulnerable to extinction. Estimates of the numbers of nonmarine molluscs vary widely, partly because many regions have not been thoroughly surveyed. There is also a shortage of specialists who can identify all the animals in any one area to species. However, in 2004 the IUCN Red List of Threatened Species included nearly 2,000 endangered nonmarine molluscs. For comparison, the great majority of mollusc species are marine, but only 41 of these appeared on the 2004 Red List. About 42% of recorded extinctions since the year 1500 are of molluscs, consisting almost entirely of nonmarine species.[28]

Hypothetical ancestral mollusc[edit] Further information: Mollusc shell Anatomical diagram of a hypothetical ancestral mollusc Because of the great range of anatomical diversity among molluscs, many textbooks start the subject of molluscan anatomy by describing what is called an archi-mollusc, hypothetical generalized mollusc, or hypothetical ancestral mollusc (HAM) to illustrate the most common features found within the phylum. The depiction is visually rather similar to modern monoplacophorans, and some suggest it may resemble very early molluscs.[7][10][13][29] The generalized mollusc is bilaterally symmetrical and has a single, "limpet-like" shell on top. The shell is secreted by a mantle covering the upper surface. The underside consists of a single muscular "foot".[10] The visceral mass, or visceropallium, is the soft, nonmuscular metabolic region of the mollusc. It contains the body organs.[8] Mantle and mantle cavity[edit] The mantle cavity, a fold in the mantle, encloses a significant amount of space. It is lined with epidermis, and is exposed, according to habitat, to sea, fresh water or air. The cavity was at the rear in the earliest molluscs, but its position now varies from group to group. The anus, a pair of osphradia (chemical sensors) in the incoming "lane", the hindmost pair of gills and the exit openings of the nephridia ("kidneys") and gonads (reproductive organs) are in the mantle cavity.[10] The whole soft body of bivalves lies within an enlarged mantle cavity.[8] Shell[edit] Main article: Mollusc shell The mantle edge secretes a shell (secondarily absent in a number of taxonomic groups, such as the nudibranchs[8]) that consists of mainly chitin and conchiolin (a protein hardened with calcium carbonate),[10][30] except the outermost layer in almost all cases is all conchiolin (see periostracum).[10] Molluscs never use phosphate to construct their hard parts,[31] with the questionable exception of Cobcrephora.[32] While most mollusc shells are composed mainly of aragonite, those gastropods that lay eggs with a hard shell use calcite (sometimes with traces of aragonite) to construct the eggshells.[33] The shell consists of three layers: the outer layer (the periostracum) made of organic matter, a middle layer made of columnar calcite, and an inner layer consisting of laminated calcite, often nacreous.[8] Foot[edit] Play media A 50-second video of snails (most likely Natica chemnitzi and Cerithium stercusmuscaram) feeding on the sea floor in the Gulf of California, Puerto Peñasco, Mexico The underside consists of a muscular foot, which has adapted to different purposes in different classes.[34]:4 The foot carries a pair of statocysts, which act as balance sensors. In gastropods, it secretes mucus as a lubricant to aid movement. In forms having only a top shell, such as limpets, the foot acts as a sucker attaching the animal to a hard surface, and the vertical muscles clamp the shell down over it; in other molluscs, the vertical muscles pull the foot and other exposed soft parts into the shell.[10] In bivalves, the foot is adapted for burrowing into the sediment;[34]:4 in cephalopods it is used for jet propulsion,[34]:4 and the tentacles and arms are derived from the foot.[35] Circulatory system[edit] Molluscs' circulatory systems are mainly open. Although molluscs are coelomates, their coeloms are reduced to fairly small spaces enclosing the heart and gonads. The main body cavity is a hemocoel through which blood and coelomic fluid circulate and which encloses most of the other internal organs. These hemocoelic spaces act as an efficient hydrostatic skeleton.[8] The blood contains the respiratory pigment hemocyanin as an oxygen-carrier. The heart consists of one or more pairs of atria (auricles), which receive oxygenated blood from the gills and pump it to the ventricle, which pumps it into the aorta (main artery), which is fairly short and opens into the hemocoel.[10] The atria of the heart also function as part of the excretory system by filtering waste products out of the blood and dumping it into the coelom as urine. A pair of nephridia ("little kidneys") to the rear of and connected to the coelom extracts any re-usable materials from the urine and dumps additional waste products into it, and then ejects it via tubes that discharge into the mantle cavity.[10] Respiration[edit] Most molluscs have only one pair of gills, or even only one gill. Generally, the gills are rather like feathers in shape, although some species have gills with filaments on only one side. They divide the mantle cavity so water enters near the bottom and exits near the top. Their filaments have three kinds of cilia, one of which drives the water current through the mantle cavity, while the other two help to keep the gills clean. If the osphradia detect noxious chemicals or possibly sediment entering the mantle cavity, the gills' cilia may stop beating until the unwelcome intrusions have ceased. Each gill has an incoming blood vessel connected to the hemocoel and an outgoing one to the heart.[10] Eating, digestion, and excretion[edit] Snail radula at work   = Food       = Radula   = Muscles   = Odontophore "belt" Members of the mollusc family use intracellular digestion to function. Most molluscs have muscular mouths with radulae, "tongues", bearing many rows of chitinous teeth, which are replaced from the rear as they wear out. The radula primarily functions to scrape bacteria and algae off rocks, and is associated with the odontophore, a cartilaginous supporting organ.[8] The radula is unique to the molluscs and has no equivalent in any other animal. Molluscs' mouths also contain glands that secrete slimy mucus, to which the food sticks. Beating cilia (tiny "hairs") drive the mucus towards the stomach, so the mucus forms a long string called a "food string".[10] At the tapered rear end of the stomach and projecting slightly into the hindgut is the prostyle, a backward-pointing cone of feces and mucus, which is rotated by further cilia so it acts as a bobbin, winding the mucus string onto itself. Before the mucus string reaches the prostyle, the acidity of the stomach makes the mucus less sticky and frees particles from it.[10] The particles are sorted by yet another group of cilia, which send the smaller particles, mainly minerals, to the prostyle so eventually they are excreted, while the larger ones, mainly food, are sent to the stomach's cecum (a pouch with no other exit) to be digested. The sorting process is by no means perfect.[10] Periodically, circular muscles at the hindgut's entrance pinch off and excrete a piece of the prostyle, preventing the prostyle from growing too large. The anus, in the part of the mantle cavity, is swept by the outgoing "lane" of the current created by the gills. Carnivorous molluscs usually have simpler digestive systems.[10] As the head has largely disappeared in bivalves, the mouth has been equipped with labial palps (two on each side of the mouth) to collect the detritus from its mucus.[8] Nervous system[edit] Simplified diagram of the mollusc nervous system The cephalic molluscs have two pairs of main nerve cords organized around a number of paired ganglia, the visceral cords serving the internal organs and the pedal ones serving the foot. Most pairs of corresponding ganglia on both sides of the body are linked by commissures (relatively large bundles of nerves). The ganglia above the gut are the cerebral, the pleural, and the visceral, which are located above the esophagus (gullet). The pedal ganglia, which control the foot, are below the esophagus and their commissure and connectives to the cerebral and pleural ganglia surround the esophagus in a circumesophageal nerve ring or nerve collar.[10] The acephalic molluscs (i.e., bivalves) also have this ring but it is less obvious and less important. The bivalves have only three pairs of ganglia— cerebral, pedal, and visceral— with the visceral as the largest and most important of the three functioning as the principal center of "thinking". Some such as the scallops have eyes around the edges of their shells which connect to a pair of looped nerves and which provide the ability to distinguish between light and shadow. Reproduction[edit] See also: Reproductive system of gastropods and Reproductive system of cephalopods Apical tuft (cilia) Prototroch (cilia) Stomach Mouth Metatroch (cilia) Mesoderm Anus /// = cilia Trochophore larva[36] The simplest molluscan reproductive system relies on external fertilization, but with more complex variations. All produce eggs, from which may emerge trochophore larvae, more complex veliger larvae, or miniature adults. Two gonads sit next to the coelom, a small cavity that surrounds the heart, into which they shed ova or sperm. The nephridia extract the gametes from the coelom and emit them into the mantle cavity. Molluscs that use such a system remain of one sex all their lives and rely on external fertilization. Some molluscs use internal fertilization and/or are hermaphrodites, functioning as both sexes; both of these methods require more complex reproductive systems.[10] The most basic molluscan larva is a trochophore, which is planktonic and feeds on floating food particles by using the two bands of cilia around its "equator" to sweep food into the mouth, which uses more cilia to drive them into the stomach, which uses further cilia to expel undigested remains through the anus. New tissue grows in the bands of mesoderm in the interior, so the apical tuft and anus are pushed further apart as the animal grows. The trochophore stage is often succeeded by a veliger stage in which the prototroch, the "equatorial" band of cilia nearest the apical tuft, develops into the velum ("veil"), a pair of cilia-bearing lobes with which the larva swims. Eventually, the larva sinks to the seafloor and metamorphoses into the adult form. While metamorphosis is the usual state in molluscs, the cephalopods differ in exhibiting direct development: the hatchling is a 'miniaturized' form of the adult.[37]

Ecology[edit] Feeding[edit] Most molluscs are herbivorous, grazing on algae or filter feeders. For those grazing, two feeding strategies are predominant. Some feed on microscopic, filamentous algae, often using their radula as a 'rake' to comb up filaments from the sea floor. Others feed on macroscopic 'plants' such as kelp, rasping the plant surface with its radula. To employ this strategy, the plant has to be large enough for the mollusc to 'sit' on, so smaller macroscopic plants are not as often eaten as their larger counterparts.[38] Filter feeders are molluscs that feed by straining suspended matter and food particle from water, typically by passing the water over their gills. Most bivalves are filter feeders. Cephalopods are primarily predatory, and the radula takes a secondary role to the jaws and tentacles in food acquisition. The monoplacophoran Neopilina uses its radula in the usual fashion, but its diet includes protists such as the xenophyophore Stannophyllum.[39] Sacoglossan sea-slugs suck the sap from algae, using their one-row radula to pierce the cell walls,[40] whereas dorid nudibranchs and some Vetigastropoda feed on sponges[41][42] and others feed on hydroids.[43] (An extensive list of molluscs with unusual feeding habits is available in the appendix of GRAHAM, A. (1955). "Molluscan diets". Journal of Molluscan Studies. 31 (3–4): 144. .)

Classification[edit] See also: List of mollusc orders Opinions vary about the number of classes of molluscs; for example, the table below shows eight living classes,[20] and two extinct ones. Although they are unlikely to form a clade, some older works combine the Caudofoveata and solenogasters into one class, the Aplacophora.[11][29] Two of the commonly recognized "classes" are known only from fossils.[18] Class Major organisms Described living species[20] Distribution Caudofoveata[11] worm-like organisms 120 seabed 200–3,000 metres (660–9,840 ft) Solenogastres[11] worm-like organisms 200 seabed 200–3,000 metres (660–9,840 ft) Polyplacophora[12] chitons 1,000 rocky tidal zone and seabed Monoplacophora[13] An ancient lineage of molluscs with cap-like shells 31 seabed 1,800–7,000 metres (5,900–23,000 ft); one species 200 metres (660 ft) Gastropoda[44] All the snails and slugs including abalone, limpets, conch, nudibranchs, sea hares, sea butterfly 70,000 marine, freshwater, land Cephalopoda[45] squid, octopus, cuttlefish, nautilus, spirula 900 marine Bivalvia[46] clams, oysters, scallops, geoducks, mussels 20,000 marine, freshwater Scaphopoda[17] tusk shells 500 marine 6–7,000 metres (20–22,966 ft) Rostroconchia †[47] fossils; probable ancestors of bivalves extinct marine Helcionelloida †[48] fossils; snail-like organisms such as Latouchella extinct marine Classification into higher taxa for these groups has been and remains problematic. A phylogenetic study suggests the Polyplacophora form a clade with a monophyletic Aplacophora.[49] Additionally, it suggests a sister taxon relationship exists between the Bivalvia and the Gastropoda. Tentaculita may also be in Mollusca (see Tentaculites). A genus called Wiwaxia probably was in the Mollusca.

Evolution[edit] Main article: Evolution of molluscs See also: Evolution of cephalopods The use of love darts by the land snail Monachoides vicinus is a form of sexual selection Fossil record[edit] Good evidence exists for the appearance of gastropods (e.g. Aldanella), cephalopods (e.g. Plectronoceras, ?Nectocaris) and bivalves (Pojetaia, Fordilla) towards the middle of the Cambrian period, c. 500 million years ago, though arguably each of these may belong only to the stem lineage of their respective classes.[50] However, the evolutionary history both of the emergence of molluscs from the ancestral group Lophotrochozoa, and of their diversification into the well-known living and fossil forms, is still vigorously debated. Debate occurs about whether some Ediacaran and Early Cambrian fossils really are molluscs. Kimberella, from about 555 million years ago, has been described by some paleontologists as "mollusc-like",[51][52] but others are unwilling to go further than "probable bilaterian".,[53][54] if that [55] There is an even sharper debate about whether Wiwaxia, from about 505 million years ago, was a mollusc, and much of this centers on whether its feeding apparatus was a type of radula or more similar to that of some polychaete worms.[53][56] Nicholas Butterfield, who opposes the idea that Wiwaxia was a mollusc, has written that earlier microfossils from 515 to 510 million years ago are fragments of a genuinely mollusc-like radula.[57] This appears to contradict the concept that the ancestral molluscan radula was mineralized.[58] The tiny Helcionellid fossil Yochelcionella is thought to be an early mollusc[48] Spirally coiled shells appear in many gastropods.[14] However, the Helcionellids, which first appear over 540 million years ago in Early Cambrian rocks from Siberia and China,[59][60] are thought to be early molluscs with rather snail-like shells. Shelled molluscs therefore predate the earliest trilobites.[48] Although most helcionellid fossils are only a few millimeters long, specimens a few centimeters long have also been found, most with more limpet-like shapes. The tiny specimens have been suggested to be juveniles and the larger ones adults.[61] Some analyses of helcionellids concluded these were the earliest gastropods.[62] However, other scientists are not convinced these Early Cambrian fossils show clear signs of the torsion that identifies modern gastropods twists the internal organs so the anus lies above the head.[14][63][64]   = Septa   = Siphuncle Septa and siphuncle in nautiloid shell Volborthella, some fossils of which predate 530 million years ago, was long thought to be a cephalopod, but discoveries of more detailed fossils showed its shell was not secreted, but built from grains of the mineral silicon dioxide (silica), and it was not divided into a series of compartments by septa as those of fossil shelled cephalopods and the living Nautilus are. Volborthella's classification is uncertain.[65] The Late Cambrian fossil Plectronoceras is now thought to be the earliest clearly cephalopod fossil, as its shell had septa and a siphuncle, a strand of tissue that Nautilus uses to remove water from compartments it has vacated as it grows, and which is also visible in fossil ammonite shells. However, Plectronoceras and other early cephalopods crept along the seafloor instead of swimming, as their shells contained a "ballast" of stony deposits on what is thought to be the underside, and had stripes and blotches on what is thought to be the upper surface.[66] All cephalopods with external shells except the nautiloids became extinct by the end of the Cretaceous period 65 million years ago.[67] However, the shell-less Coleoidea (squid, octopus, cuttlefish) are abundant today.[68] The Early Cambrian fossils Fordilla and Pojetaia are regarded as bivalves.[69][70][71][72] "Modern-looking" bivalves appeared in the Ordovician period, 488 to 443 million years ago.[73] One bivalve group, the rudists, became major reef-builders in the Cretaceous, but became extinct in the Cretaceous–Paleogene extinction event.[74] Even so, bivalves remain abundant and diverse. The Hyolitha are a class of extinct animals with a shell and operculum that may be molluscs. Authors who suggest they deserve their own phylum do not comment on the position of this phylum in the tree of life[75] Phylogeny[edit] Lophotrochozoa Brachiopods Mollusca Bivalves Monoplacophorans ("limpet-like", "living fossils") Gastropods (snails, slugs, limpets, sea hares) Cephalopods (nautiloids, ammonites, squid, etc.) Scaphopods (tusk shells) Aplacophorans (spicule-covered, worm-like) Polyplacophorans (chitons) Halwaxiids Wiwaxia Halkieria Orthrozanclus Odontogriphus A possible "family tree" of molluscs (2007).[76][77] Does not include annelid worms as the analysis concentrated on fossilizable "hard" features.[76] The phylogeny (evolutionary "family tree") of molluscs is a controversial subject. In addition to the debates about whether Kimberella and any of the "halwaxiids" were molluscs or closely related to molluscs,[52][53][56][57] debates arise about the relationships between the classes of living molluscs.[54] In fact, some groups traditionally classified as molluscs may have to be redefined as distinct but related.[78] Molluscs are generally regarded members of the Lophotrochozoa,[76] a group defined by having trochophore larvae and, in the case of living Lophophorata, a feeding structure called a lophophore. The other members of the Lophotrochozoa are the annelid worms and seven marine phyla.[79] The diagram on the right summarizes a phylogeny presented in 2007. Because the relationships between the members of the family tree are uncertain, it is difficult to identify the features inherited from the last common ancestor of all molluscs.[80] For example, it is uncertain whether the ancestral mollusc was metameric (composed of repeating units)—if it was, that would suggest an origin from an annelid-like worm.[81] Scientists disagree about this: Giribet and colleagues concluded, in 2006, the repetition of gills and of the foot's retractor muscles were later developments,[7] while in 2007, Sigwart concluded the ancestral mollusc was metameric, and it had a foot used for creeping and a "shell" that was mineralized.[54] In one particular branch of the family tree, the shell of conchiferans is thought to have evolved from the spicules (small spines) of aplacophorans; but this is difficult to reconcile with the embryological origins of spicules.[80] The molluscan shell appears to have originated from a mucus coating, which eventually stiffened into a cuticle. This would have been impermeable and thus forced the development of more sophisticated respiratory apparatus in the form of gills.[48] Eventually, the cuticle would have become mineralized,[48] using the same genetic machinery (engrailed) as most other bilaterian skeletons.[81] The first mollusc shell almost certainly was reinforced with the mineral aragonite.[30] The evolutionary relationships 'within' the molluscs are also debated, and the diagrams below show two widely supported reconstructions: Molluscs Aculifera Solenogastres Caudofoveata Polyplacophorans Conchifera Monoplacophorans Bivalves Scaphopods Gastropods Cephalopods The "Aculifera" hypothesis[76] Molluscs Solenogastres Caudofoveata Testaria Polyplacophorans Monoplacophorans Bivalves Scaphopods Gastropods Cephalopods The "Testaria" hypothesis[76] Morphological analyses tend to recover a conchiferan clade that receives less support from molecular analyses,[82] although these results also lead to unexpected paraphylies, for instance scattering the bivalves throughout all other mollusc groups.[83] However, an analysis in 2009 using both morphological and molecular phylogenetics comparisons concluded the molluscs are not monophyletic; in particular, Scaphopoda and Bivalvia are both separate, monophyletic lineages unrelated to the remaining molluscan classes; the traditional phylum Mollusca is polyphyletic, and it can only be made monophyletic if scaphopods and bivalves are excluded.[78] A 2010 analysis recovered the traditional conchiferan and aculiferan groups, and showed molluscs were monophyletic, demonstrating that available data for solenogastres was contaminated.[84] Current molecular data are insufficient to constrain the molluscan phylogeny, and since the methods used to determine the confidence in clades are prone to overestimation, it is risky to place too much emphasis even on the areas of which different studies agree.[85] Rather than eliminating unlikely relationships, the latest studies add new permutations of internal molluscan relationships, even bringing the conchiferan hypothesis into question.[86]

Human interaction[edit] Main article: Molluscs in culture For millennia, molluscs have been a source of food for humans, as well as important luxury goods, notably pearls, mother of pearl, Tyrian purple dye, sea silk, and chemical compounds. Their shells have also been used as a form of currency in some preindustrial societies. A number of species of molluscs can bite or sting humans, and some have become agricultural pests. Uses by humans[edit] Further information: Seashell and List of edible molluscs Molluscs, especially bivalves such as clams and mussels, have been an important food source since at least the advent of anatomically modern humans, and this has often resulted in overfishing.[87] Other commonly eaten molluscs include octopuses and squids, whelks, oysters, and scallops.[88] In 2005, China accounted for 80% of the global mollusc catch, netting almost 11,000,000 tonnes (11,000,000 long tons; 12,000,000 short tons). Within Europe, France remained the industry leader.[89] Some countries regulate importation and handling of molluscs and other seafood, mainly to minimize the poison risk from toxins that can sometimes accumulate in the animals.[90] Saltwater pearl oyster farm in Seram, Indonesia Most molluscs with shells can produce pearls, but only the pearls of bivalves and some gastropods, whose shells are lined with nacre, are valuable.[14][16] The best natural pearls are produced by marine pearl oysters, Pinctada margaritifera and Pinctada mertensi, which live in the tropical and subtropical waters of the Pacific Ocean. Natural pearls form when a small foreign object gets stuck between the mantle and shell. The two methods of culturing pearls insert either "seeds" or beads into oysters. The "seed" method uses grains of ground shell from freshwater mussels, and overharvesting for this purpose has endangered several freshwater mussel species in the southeastern United States.[16] The pearl industry is so important in some areas, significant sums of money are spent on monitoring the health of farmed molluscs.[91] Byzantine Emperor Justinian I clad in Tyrian purple and wearing numerous pearls Other luxury and high-status products were made from molluscs. Tyrian purple, made from the ink glands of murex shells, "... fetched its weight in silver" in the fourth century BC, according to Theopompus.[92] The discovery of large numbers of Murex shells on Crete suggests the Minoans may have pioneered the extraction of "imperial purple" during the Middle Minoan period in the 20th–18th centuries BC, centuries before the Tyrians.[93][94] Sea silk is a fine, rare, and valuable fabric produced from the long silky threads (byssus) secreted by several bivalve molluscs, particularly Pinna nobilis, to attach themselves to the sea bed.[95] Procopius, writing on the Persian wars circa 550 CE, "stated that the five hereditary satraps (governors) of Armenia who received their insignia from the Roman Emperor were given chlamys (or cloaks) made from lana pinna. Apparently, only the ruling classes were allowed to wear these chlamys."[96] Mollusc shells, including those of cowries, were used as a kind of money (shell money) in several preindustrial societies. However, these "currencies" generally differed in important ways from the standardized government-backed and -controlled money familiar to industrial societies. Some shell "currencies" were not used for commercial transactions, but mainly as social status displays at important occasions, such as weddings.[97] When used for commercial transactions, they functioned as commodity money, as a tradable commodity whose value differed from place to place, often as a result of difficulties in transport, and which was vulnerable to incurable inflation if more efficient transport or "goldrush" behavior appeared.[98] Bioindicators[edit] Bivalve molluscs are used as bioindicators to monitor the health of aquatic environments in both fresh water and the marine environments. Their population status or structure, physiology, behaviour or the level of contamination with elements or compounds can indicate the state of contamination status of the ecosystem. They are particularly useful since they are sessile so that they are representative of the environment where they are sampled or placed.[99] Potamopyrgus antipodarum is used by some water treatment plants to test for estrogen-mimicking pollutants from industrial agriculture. Harmful to humans[edit] Stings and bites[edit] The blue-ringed octopus's rings are a warning signal; this octopus is alarmed, and its bite can kill.[100] Some molluscs sting or bite, but deaths from mollusc venoms total less than 10% of those from jellyfish stings.[101] All octopuses are venomous,[102] but only a few species pose a significant threat to humans. Blue-ringed octopuses in the genus Hapalochlaena, which live around Australia and New Guinea, bite humans only if severely provoked,[100] but their venom kills 25% of human victims. Another tropical species, Octopus apollyon, causes severe inflammation that can last for over a month even if treated correctly,[103] and the bite of Octopus rubescens can cause necrosis that lasts longer than one month if untreated, and headaches and weakness persisting for up to a week even if treated.[104] Live cone snails can be dangerous to shell collectors, but are useful to neurology researchers.[105] All species of cone snails are venomous and can sting painfully when handled, although many species are too small to pose much of a risk to humans, and only a few fatalities have been reliably reported. Their venom is a complex mixture of toxins, some fast-acting and others slower but deadlier.[105][101][106] The effects of individual cone-shell toxins on victims' nervous systems are so precise as to be useful tools for research in neurology, and the small size of their molecules makes it easy to synthesize them.[105][107] Disease vectors[edit] Skin vesicles created by the penetration of Schistosoma. (Source: CDC) Schistosomiasis (also known as bilharzia, bilharziosis or snail fever), a disease caused by the fluke worm Schistosoma, is "second only to malaria as the most devastating parasitic disease in tropical countries. An estimated 200 million people in 74 countries are infected with the disease – 100 million in Africa alone."[108] The parasite has 13 known species, two of which infect humans. The parasite itself is not a mollusc, but all the species have freshwater snails as intermediate hosts.[109] Pests[edit] Some species of molluscs, particularly certain snails and slugs, can be serious crop pests,[110] and when introduced into new environments, can unbalance local ecosystems. One such pest, the giant African snail Achatina fulica, has been introduced to many parts of Asia, as well as to many islands in the Indian Ocean and Pacific Ocean. In the 1990s, this species reached the West Indies. Attempts to control it by introducing the predatory snail Euglandina rosea proved disastrous, as the predator ignored Achatina fulica and went on to extirpate several native snail species, instead.[111]

Notes[edit] ^ The formerly dominant spelling mollusk is still used in the U.S. — see the reasons given in Gary Rosenberg's "Mollusckque - Mollusk vs Mollusc". For the spelling mollusc, see the reasons given in: Brusca & Brusca. Invertebrates (2nd ed.). 

References[edit] ^ a b c d Chapman, A.D. (2009). Numbers of Living Species in Australia and the World, 2nd edition. Australian Biological Resources Study, Canberra. Retrieved 12 January 2010. ISBN 978-0-642-56860-1 (printed); ISBN 978-0-642-56861-8 (online). ^ Rosenberg, G. (2014). A new critical estimate of named species-level diversity of the recent Mollusca. American Malacological Bulletin, 32(2), 308-322. ^ Taylor, P.D., & Lewis, D.N. (2005). Fossil invertebrates. Harvard University Press, 208 pp. ^ Little, L.; Fowler, H.W.; Coulson, J.; Onions, C.T., eds. (1964). "Malacology". Shorter Oxford English Dictionary. Oxford University press.  ^  Chisholm, Hugh, ed. (1911). "Molluscoida". Encyclopædia Britannica. 18 (11th ed.). Cambridge University Press. p. 675.  ^ Hogan, C. Michael. (2010). "Calcium". In Jorgensen, A.; Cleveland, C. Encyclopedia of Earth. National Council for Science and the Environment.  ^ a b c d Giribet, G.; Okusu, A.; Lindgren, A.R.; Huff, S.W.; Schrödl, M.; Nishiguchi, M.K. (May 2006). "Evidence for a clade composed of molluscs with serially repeated structures: monoplacophorans are related to chitons" (Free full text). Proceedings of the National Academy of Sciences of the United States of America. 103 (20): 7723–7728. Bibcode:2006PNAS..103.7723G. doi:10.1073/pnas.0602578103. PMC 1472512 . PMID 16675549.  ^ a b c d e f g h Hayward, PJ (1996). Handbook of the Marine Fauna of North-West Europe. Oxford University Press. pp. 484–628. ISBN 0-19-854055-8.  ^ a b c Brusca, R.C. & Brusca, G.J. (2003). Invertebrates (2 ed.). Sinauer Associates. p. 702. ISBN 0-87893-097-3.  ^ a b c d e f g h i j k l m n o p q Ruppert, pp. 284–291 ^ a b c d Ruppert, pp. 291–292 ^ a b Ruppert, pp. 292–298 ^ a b c Ruppert, pp. 298–300 ^ a b c d Ruppert, pp. 300–343 ^ Ruppert, pp. 343–367 ^ a b c Ruppert, pp. 367–403 ^ a b Ruppert, pp. 403–407 ^ a b c d Ponder, W.F.; Lindberg, D.R., eds. (2008). Phylogeny and Evolution of the Mollusca. Berkeley: University of California Press. p. 481. ISBN 978-0-520-25092-5.  ^ Nicol, David (June 1969). "The Number of Living Species of Molluscs". Systematic Zoology. 18 (2): 251–254. doi:10.2307/2412618. JSTOR 2412618.  ^ a b c Haszprunar, G. (2001). "Mollusca (Molluscs)". Encyclopedia of Life Sciences. John Wiley & Sons, Ltd. doi:10.1038/npg.els.0001598.  ^ Hancock, Rebecca (2008). "Recognising research on molluscs". Australian Museum. Retrieved 2009-03-09.  ^ Ruppert, Front endpaper 1 ^ Ponder, Winston F. & Lindberg, David R. (2004). "Phylogeny of the Molluscs". World Congress of Malacology. Retrieved 2009-03-09.  ^ Raup, David M. & Stanley, Steven M. (1978). Principles of Paleontology (2 ed.). W.H. Freeman and Co. pp. 4–5. ISBN 0716700220.  ^ Barnes, R.S.K.; Calow, P.; Olive, P.J.W.; Golding, D.W.; Spicer, J.I. (2001). The Invertebrates, A Synthesis (3 ed.). UK: Blackwell Science.  ^ Kubodera, T.; Mori, K. (December 22, 2005). "First-ever observations of a live giant squid in the wild" (PDF). Proceedings of the Royal Society B. 272 (1581): 2583–2586. doi:10.1098/rspb.2005.3158. PMC 1559985 . PMID 16321779. Retrieved 2008-10-22.  ^ Black, Richard (April 26, 2008). "Colossal squid out of the freezer". BBC News. Retrieved 2008-10-01.  ^ Lydeard, C.; Cowie, R.; Ponder, W.F.; et al. (April 2004). "The global decline of nonmarine mollusks" (PDF). BioScience. 54 (4): 321–330. doi:10.1641/0006-3568(2004)054[0321:TGDONM]2.0.CO;2. Archived from the original on March 31, 2007. CS1 maint: Date and year (link) CS1 maint: Unfit url (link) ^ a b Healy, J.M. (2001). "The Mollusca". In Anderson, D.T. Invertebrate Zoology (2 ed.). Oxford University Press. pp. 120–171. ISBN 0-19-551368-1.  ^ a b Porter, S. (June 1, 2007). "Seawater Chemistry and Early Carbonate Biomineralization". Science. 316 (5829): 1302. Bibcode:2007Sci...316.1302P. doi:10.1126/science.1137284. PMID 17540895.  ^ Yochelson, E. L. (1975). "Discussion of early Cambrian "molluscs"" (PDF). Journal of the Geological Society. 131 (6): 661–662. doi:10.1144/gsjgs.131.6.0661.  ^ Cherns, L. (December 2004). "Early Palaeozoic diversification of chitons (Polyplacophora, Mollusca) based on new data from the Silurian of Gotland, Sweden". Lethaia. 37 (4): 445–456. doi:10.1080/00241160410002180.  ^ Tompa, A. S. (December 1976). "A comparative study of the ultrastructure and mineralogy of calcified land snail eggs (Pulmonata: Stylommatophora)". Journal of Morphology. 150 (4): 861–887. doi:10.1002/jmor.1051500406. hdl:2027.42/50263.  ^ a b c Wilbur, Karl M.; Trueman, E.R.; Clarke, M.R., eds. (1985), The Mollusca, 11. Form and Function, New York: Academic Press, ISBN 0-12-728702-7  ^ Shigeno, S; Sasaki, T; Moritaki, T; Kasugai, T; Vecchione, M; Agata, K (Jan 2008). "Evolution of the cephalopod head complex by assembly of multiple molluscan body parts: Evidence from Nautilus embryonic development". Journal of Morphology. 269 (1): 1–17. doi:10.1002/jmor.10564. PMID 17654542.  ^ Ruppert, E.E.; Fox, R.S. & Barnes, R.D. (2004). "Mollusca". Invertebrate Zoology (7th ed.). Brooks / Cole. pp. 290–291. ISBN 0030259827.  ^ Marin, F.; Luquet, G. (October 2004). "Molluscan shell proteins". Comptes Rendus Palevol. 3 (6–7): 469. doi:10.1016/j.crpv.2004.07.009.  ^ Steneck, R.S.; Watling, L. (July 1982). "Feeding capabilities and limitation of herbivorous molluscs: A functional group approach". Marine Biology. 68 (3): 299–319. doi:10.1007/BF00409596.  ^ Tendal O.S. (1985). "Xenophyophores (Protozoa, Sarcodina) in the diet of Neopilina galatheae (Mollusca, Monoplacophora)" (PDF). Galathea Report. 16: 95–98. Retrieved 2013-09-14.  ^ Jensen, K. R. (February 1993). "Morphological adaptations and plasticity of radular teeth of the Sacoglossa (= Ascoglossa) (Mollusca: Opisthobranchia) in relation to their food plants". Biological Journal of the Linnean Society. 48 (2): 135–155. doi:10.1111/j.1095-8312.1993.tb00883.x.  ^ Wägele, H. (March 1989). "Diet of some Antarctic nudibranchs (Gastropoda, Opisthobranchia, Nudibranchia)". Marine Biology. 100 (4): 439–441. doi:10.1007/BF00394819.  ^ Publishers, Bentham Science (July 1999). Current Organic Chemistry.  ^ Lambert, W. J. (1991-10-01). "Coexistence of Hydroid Eating Nudibranchs: Do Feeding Biology and Habitat Use Matter?".  Missing or empty |url= (help) ^ Ruppert, p. 300 ^ Ruppert, p. 343 ^ Ruppert, p. 367 ^ Clarkson, E.N.K. (1998). Invertebrate Palaeontology and Evolution. Blackwell. p. 221. ISBN 0-632-05238-4.  ^ a b c d e Runnegar, B.; Pojeta Jr, J. (October 1974). "Molluscan Phylogeny: the Paleontological Viewpoint". Science. 186 (4161): 311–317. Bibcode:1974Sci...186..311R. doi:10.1126/science.186.4161.311. JSTOR 1739764. PMID 17839855.  ^ Kocot, K. M.; Cannon, J. T.; Todt, C.; Citarella, M. R.; Kohn, A. B.; Meyer, A.; Santos, S. R.; Schander, C.; Moroz, L. L.; et al. (September 22, 2011). "Phylogenomics reveals deep molluscan relationships". Nature. 477 (7365): 452–456. Bibcode:2011Natur.477..452K. doi:10.1038/nature10382. PMID 21892190.  ^ Budd, G. E. & Jensen, S. A critical reappraisal of the fossil record of the bilaterian phyla. Biol. Rev. 75, 253–295 (2000). ^ Fedonkin, M.A.; Waggoner, B.M. (August 28, 1997). "The Late Precambrian fossil Kimberella is a mollusc-like bilaterian organism". Nature. 388 (6645): 868. Bibcode:1997Natur.388..868F. doi:10.1038/42242.  ^ a b Fedonkin, M.A.; Simonetta, A.; Ivantsov, A.Y. (2007). "New data on Kimberella, the Vendian mollusc-like organism (White Sea region, Russia): palaeoecological and evolutionary implications" (PDF). Geological Society, London, Special Publications. 286: 157–179. Bibcode:2007GSLSP.286..157F. doi:10.1144/SP286.12. Retrieved 2008-07-10.  ^ a b c Butterfield, N.J. (2006). "Hooking some stem-group "worms": fossil lophotrochozoans in the Burgess Shale". BioEssays. 28 (12): 1161–6. doi:10.1002/bies.20507. PMID 17120226.  ^ a b c Sigwart, J. D.; Sutton, M. D. (October 2007). "Deep molluscan phylogeny: synthesis of palaeontological and neontological data". Proceedings of the Royal Society B: Biological Sciences. 274 (1624): 2413–2419. doi:10.1098/rspb.2007.0701. PMC 2274978 . PMID 17652065.  For a summary, see "The Mollusca". University of California Museum of Paleontology. Retrieved 2 October 2008.  ^ Budd, G. E., and S. Jensen. 2016: The origin of the animals and a “Savannah” hypothesis for early bilaterian evolution. Biological Reviews 7:Online ahead of print. ^ a b Caron, J.B.; Scheltema, A.; Schander, C.; Rudkin, D. (July 13, 2006). "A soft-bodied mollusc with radula from the Middle Cambrian Burgess Shale". Nature. 442 (7099): 159–163. Bibcode:2006Natur.442..159C. doi:10.1038/nature04894. PMID 16838013.  ^ a b Butterfield, N.J. (May 2008). "An Early Cambrian Radula". Journal of Paleontology. 82 (3): 543–554. doi:10.1666/07-066.1.  ^ Cruz, R.; Lins, U.; Farina, M. (1998). "Minerals of the radular apparatus of Falcidens sp. (Caudofoveata) and the evolutionary implications for the Phylum Mollusca". Biological Bulletin. 194 (2): 224–230. doi:10.2307/1543051. JSTOR 1543051. PMID 28570844.  ^ Parkhaev, P. Yu. (2007). "The Cambrian 'basement' of gastropod evolution". Geological Society, London, Special Publications. 286: 415–421. Bibcode:2007GSLSP.286..415P. doi:10.1144/SP286.31. ISBN 978-1-86239-233-5. Retrieved 2009-11-01.  ^ Steiner, M.; Li, G.; Qian, Y.; Zhu, M.; Erdtmann, B.D. (2007). "Neoproterozoic to Early Cambrian small shelly fossil assemblages and a revised biostratigraphic correlation of the Yangtze Platform (China)". Palaeogeography, Palaeoclimatology, Palaeoecology. 254: 67. doi:10.1016/j.palaeo.2007.03.046.  ^ Mus, M.M.; Palacios, T.; Jensen, S. (2008). "Size of the earliest mollusks: Did small helcionellids grow to become large adults?". Geology. 36 (2): 175. Bibcode:2008Geo....36..175M. doi:10.1130/G24218A.1.  ^ Landing, E.; Geyer, G.; Bartowski, K. E. (2002). "Latest Early Cambrian Small Shelly Fossils, Trilobites, and Hatch Hill Dysaerobic Interval on the Quebec Continental Slope". Journal of Paleontology. 76 (2): 287–305. doi:10.1666/0022-3360(2002)076<0287:LECSSF>2.0.CO;2. JSTOR 1307143. CS1 maint: Date and year (link) ^ Frýda, J.; Nützel, A.; Wagner, P.J. (2008). "Paleozoic Gastropoda". In Ponder, W.F.; Lindberg, D.R. Phylogeny and evolution of the Mollusca. California Press. pp. 239–264. ISBN 0-520-25092-3.  ^ Kouchinsky, A. (2000). "Shell microstructures in Early Cambrian molluscs" (PDF). Acta Palaeontologica Polonica. 45 (2): 119–150. Retrieved 4 Nov 2009.  ^ Hagadorn, J.W. & Waggoner, B.M. (2002). "The Early Cambrian problematic fossil Volborthella: New insights from the Basin and Range". In Corsetti, F.A. Proterozoic-Cambrian of the Great Basin and Beyond, Pacific Section SEPM Book 93 (PDF). SEPM (Society for Sedimentary Geology). pp. 135–150. Archived from the original on 2006-09-11. CS1 maint: BOT: original-url status unknown (link) ^ Vickers-Rich, P.; Fenton, C.L.; Fenton, M.A.; Rich, T.H. (1997). The Fossil Book: A Record of Prehistoric Life. Courier Dover Publications. pp. 269–272. ISBN 0-486-29371-8.  ^ Marshall C.R.; Ward P.D. (1996). "Sudden and Gradual Molluscan Extinctions in the Latest Cretaceous of Western European Tethys". Science. 274 (5291): 1360–1363. Bibcode:1996Sci...274.1360M. doi:10.1126/science.274.5291.1360. PMID 8910273.  ^ Monks, N. "A Broad Brush History of the Cephalopoda". Retrieved 2009-03-21.  ^ Pojeta, J. (2000). "Cambran Pelecypoda (Mollusca)". American Malacological Bulletin. 15: 157–166.  ^ Schneider, J.A. (2001). "Bivalve systematics during the 20th century". Journal of Paleontology. 75 (6): 1119–1127. doi:10.1666/0022-3360(2001)075<1119:BSDTC>2.0.CO;2. CS1 maint: Date and year (link) ^ Gubanov, A.P.; Kouchinsky, A.V.; Peel, J.S. (2007). "The first evolutionary-adaptive lineage within fossil molluscs". Lethaia. 32 (2): 155. doi:10.1111/j.1502-3931.1999.tb00534.x.  ^ Gubanov, A.P.; Peel, J.S. (2003). "The early Cambrian helcionelloid mollusc Anabarella Vostokova". Palaeontology. 46 (5): 1073–1087. doi:10.1111/1475-4983.00334.  ^ Zong-Jie, F. (2006). "An introduction to Ordovician bivalves of southern China, with a discussion of the early evolution of the Bivalvia". Geological Journal. 41 (3–4): 303–328. doi:10.1002/gj.1048.  ^ Raup, D.M.; Jablonski, D. (1993). "Geography of end-Cretaceous marine bivalve extinctions". Science. 260 (5110): 971–973. Bibcode:1993Sci...260..971R. doi:10.1126/science.11537491. PMID 11537491.  ^ Malinky, J.M. (2009). "Permian Hyolithida from Australia: The Last of the Hyoliths?". Journal of Paleontology. 83: 147–152. doi:10.1666/08-094R.1.  ^ a b c d e Sigwart, J.D.; Sutton, M.D. (October 2007). "Deep molluscan phylogeny: synthesis of palaeontological and neontological data". Proceedings of the Royal Society B. 274 (1624): 2413–2419. doi:10.1098/rspb.2007.0701. PMC 2274978 . PMID 17652065.  For a summary, see "The Mollusca". University of California Museum of Paleontology. Retrieved 2008-10-02.  ^ "The Mollusca". University of California Museum of Paleontology. Retrieved 2008-10-02.  ^ a b Goloboff, Pablo A.; Catalano, Santiago A.; Mirande, J. Marcos; Szumik, Claudia A.; Arias, J. Salvador; Källersjö, Mari; Farris, James S. (2009). "Phylogenetic analysis of 73 060 taxa corroborates major eukaryotic groups". Cladistics. 25 (3): 211–230. doi:10.1111/j.1096-0031.2009.00255.x.  ^ "Introduction to the Lophotrochozoa". University of California Museum of Paleontology. Retrieved 2008-10-02.  ^ a b Henry, J.; Okusu, A.; Martindale, M. (2004). "The cell lineage of the polyplacophoran, Chaetopleura apiculata: variation in the spiralian program and implications for molluscan evolution". Developmental Biology. 272 (1): 145–160. doi:10.1016/j.ydbio.2004.04.027. PMID 15242797.  ^ a b Jacobs, D.K.; Wray, C. G.; Wedeen, C. J.; Kostriken, R.; Desalle, R.; Staton, J. L.; Gates, R.D.; Lindberg, D.R. (2000). "Molluscan engrailed expression, serial organization, and shell evolution". Evolution & Development. 2 (6): 340–347. doi:10.1046/j.1525-142x.2000.00077.x. PMID 11256378.  ^ Winnepenninckx, B; Backeljau, T; De Wachter, R (1996). "Investigation of molluscan phylogeny on the basis of 18S rRNA sequences". Molecular Biology and Evolution. 13 (10): 1306–1317. doi:10.1093/oxfordjournals.molbev.a025577. PMID 8952075.  ^ Passamaneck, Y.; Schander, C.; Halanych, K. (2004). "Investigation of molluscan phylogeny using large-subunit and small-subunit nuclear rRNA sequences". Molecular Phylogenetics & Evolution. 32 (1): 25–38. doi:10.1016/j.ympev.2003.12.016. PMID 15186794.  ^ Wilson, N.; Rouse, G.; Giribet, G. (2010). "Assessing the molluscan hypothesis Serialia (Monoplacophora+Polyplacophora) using novel molecular data". Molecular Phylogenetics & Evolution. 54 (1): 187–193. doi:10.1016/j.ympev.2009.07.028. PMID 19647088.  ^ Wägele, J.; Letsch, H.; Klussmann-Kolb, A.; Mayer, C.; Misof, B.; Wägele, H. (2009). "Phylogenetic support values are not necessarily informative: the case of the Serialia hypothesis (a mollusk phylogeny)". Frontiers in Zoology. 6 (1): 12. doi:10.1186/1742-9994-6-12. PMC 2710323 . PMID 19555513.  ^ Vinther, J.; Sperling, E. A.; Briggs, D. E. G.; Peterson, K. J. (2011). "A molecular palaeobiological hypothesis for the origin of aplacophoran molluscs and their derivation from chiton-like ancestors". Proceedings of the Royal Society B: Biological Sciences. 279 (1732): 1259. doi:10.1098/rspb.2011.1773. PMC 3282371 . PMID 21976685.  ^ Mannino, M.A.; Thomas, K.D. (2002). "Depletion of a resource? The impact of prehistoric human foraging on intertidal mollusc communities and its significance for human settlement, mobility and dispersal". World Archaeology. 33 (3): 452–474. doi:10.1080/00438240120107477. JSTOR 827879.  ^ Garrow, J.S.; Ralph, A.; James, W.P.T. (2000). Human Nutrition and Dietetics. Elsevier Health Sciences. p. 370. ISBN 0-443-05627-7.  ^ "China catches almost 11 m tonnes of molluscs in 2005". FAO. Retrieved 2008-10-03.  ^ "Importing fishery products or bivalve molluscs". United Kingdom: Food Standards Agency. Retrieved 2008-10-02.  ^ Jones, J.B.; Creeper, J. (April 2006). "Diseases of Pearl Oysters and Other Molluscs: a Western Australian Perspective". Journal of Shellfish Research. 25 (1): 233–238. doi:10.2983/0730-8000(2006)25[233:DOPOAO]2.0.CO;2. CS1 maint: Date and year (link) ^ The fourth-century BC historian Theopompus, cited by Athenaeus (12:526) around 200 BC ; according to Gulick, C.B. (1941). Athenaeus, The Deipnosophists. Cambridge, Mass.: Harvard University Press. ISBN 0-674-99380-2.  ^ Reese, D.S. (1987). "Palaikastro Shells and Bronze Age Purple-Dye Production in the Mediterranean Basin". Annual of the British School of Archaeology at Athens. 82: 201–6. doi:10.1017/s0068245400020438.  ^ Stieglitz, R.R. (March 1994). "The Minoan Origin of Tyrian Purple". Biblical Archaeologist. 57 (1): 46–54. doi:10.2307/3210395. JSTOR 3210395.  ^ Webster's Third New International Dictionary (Unabridged) 1976. G. & C. Merriam Co., p. 307. ^ Turner, R.D.; Rosewater, J. (June 1958). "The Family Pinnidae in the Western Atlantic". Johnsonia. 3 (38): 294.  ^ Maurer, B. (October 2006). "The Anthropology of Money" (PDF). Annual Review of Anthropology. 35: 15–36. doi:10.1146/annurev.anthro.35.081705.123127. [dead link] ^ Hogendorn, J. & Johnson, M. (2003). The Shell Money of the Slave Trade. Cambridge University Press. ISBN 0521541107.  Particularly chapters "Boom and slump for the cowrie trade" (pages 64–79) and "The cowrie as money: transport costs, values and inflation" (pages 125–147) ^ Université Bordeaux; et al. "MolluSCAN eye project". Retrieved 28 January 2017. CS1 maint: Explicit use of et al. (link) ^ a b Alafaci, A. "Blue ringed octopus". Australian Venom Research Unit. Retrieved 2008-10-03.  ^ a b Williamson, J.A.; Fenner, P.J.; Burnett, J.W.; Rifkin, J. (1996). Venomous and Poisonous Marine Animals: A Medical and Biological Handbook. UNSW Press. pp. 65–68. ISBN 0-86840-279-6.  ^ Anderson, R.C. (1995). "Aquarium husbandry of the giant Pacific octopus". Drum and Croaker. 26: 14–23.  ^ Brazzelli, V.; Baldini, F.; Nolli, G.; Borghini, F.; Borroni, G. (March 1999). "Octopus apollyon bite". Contact Dermatitis. 40 (3): 169. doi:10.1111/j.1600-0536.1999.tb06025.x. PMID 10073455.  ^ Anderson, R.C. (1999). "An octopus bite and its treatment". The Festivus. 31: 45–46.  ^ a b c Concar, D. (19 October 1996). "Doctor snail—Lethal to fish and sometimes even humans, cone snail venom contains a pharmacopoeia of precision drugs". New Scientist. Retrieved 2008-10-03.  ^ Livett, B. "Cone Shell Mollusc Poisoning, with Report of a Fatal Case". Department of Biochemistry and Molecular Biology, University of Melbourne.  ^ Haddad Junior, V.; Paula Neto, J.O.B.D.; Cobo, V.L.J. (September–October 2006). "Venomous mollusks: The risks of human accidents by conus snails (gastropoda: Conidae) in Brazil". Revista da Sociedade Brasileira de Medicina Tropical. 39 (5): 498. doi:10.1590/S0037-86822006000500015. PMID 17160331.  ^ "The Carter Center Schistosomiasis Control Program". The Carter Center. Retrieved 2008-10-03.  ^ Brown, D.S. (1994). Freshwater Snails of Africa and Their Medical Importance. CRC Press. p. 305. ISBN 0-7484-0026-5.  ^ Barker, G.M. (2002). Molluscs As Crop Pests. CABI Publications. ISBN 0-85199-320-6.  ^ Civeyrel, L.; Simberloff, D. (October 1996). "A tale of two snails: is the cure worse than the disease?". Biodiversity and Conservation. 5 (10): 1231–1252. doi:10.1007/BF00051574. 

Bibliography[edit] Ruppert, E.E.; Fox, R.S.; Barnes, R.D. (2004). Invertebrate Zoology (7 ed.). Brooks / Cole. ISBN 0-03-025982-7. 

Further reading[edit] Sturm, C., T.A. Pearce, and A. Valdes. The Mollusks: A Guide to their Study, Collection, and Preservation. Universal Publishers. 2006. 454 pages. ISBN 1581129300

External links[edit] Wikimedia Commons has media related to Mollusca. The Wikibook Dichotomous Key has a page on the topic of: Mollusca Data related to Mollusca at Wikispecies Mollusca at the Encyclopedia of Life Researchers complete mollusk evolutionary tree; 26 October 2011 Hardy's Internet Guide to Marine Gastropods Rotterdam Natural History Museum Shell Image Gallery Mussel Watch Programme Online biomonitoring of bivalve activity, 24/7: MolluSCAN eye v t e Extant Animal phyla Domain Archaea Bacteria Eukaryota (Supergroup Plant Hacrobia Heterokont Alveolata Rhizaria Excavata Amoebozoa Opisthokonta Animal Fungi) A n i m a l i a Hexactinellida (glass sponges) D+C+H+E Demospongiae (demosponges) C+H+E Calcarea (calcareous sponges) H+E Homoscleromorpha Diploblasts (Eumetazoa) Ctenophora (comb jellies) ParaHoxozoa Placozoa (Trichoplax) Planulozoa Cnidaria (jellyfish and relatives) Bilateria (Triploblasts) (see below↓) Bilateria Xenacoelomorpha Xenoturbellida (Xenoturbella) Acoelomorpha acoels nemertodermatids N e p h r o z o a Deuterostomia Chordata lancelets tunicates craniates / vertebrates Ambulacraria Echinodermata (starfish and relatives) Hemichordata acorn worms pterobranchs P r o t o s t o m i a Ecdysozoa Scalidophora Kinorhyncha (mud dragons) Priapulida (penis worms) N+L+P Nematoida Nematoda (roundworms) Nematomorpha (horsehair worms) L+P Loricifera Panarthropoda Onychophora (velvet worms) Tactopoda Arthropoda (arthropods) Tardigrada (waterbears) S p i r a l i a Gnathifera¹ Chaetognatha (arrow worms) Gnathostomulida (jaw worms) Micrognathozoa (Limnognathia) Syndermata Rotifera Acanthocephala Platytrochozoa R+M Mesozoa Orthonectida Dicyemida or Rhombozoa Rouphozoa¹ Platyhelminthes (flatworms) Gastrotricha (hairybacks) Lophotrochozoa Cycliophora (Symbion) Mollusca (molluscs) A+N Annelida (ringed worms) Nemertea (ribbon worms) Lophophorata Bryozoa Entoprocta or Kamptozoa Ectoprocta (moss animals) Brachiozoa Brachiopoda (lamp shells) Phoronida (horseshoe worms) Major groups within phyla Sponges Calcareous Hexactinellid Demosponge Homoscleromorpha Cnidarians Anthozoa inc. corals Medusozoa inc. jellyfish Myxozoa Vertebrates Jawless fish Cartilaginous fish Bony fish Amphibians Reptiles/Birds Mammals Echinoderms Sea lilies Asterozoa inc. starfish Echinozoa Nematodes Chromadorea Enoplea Secernentea Arthropods Chelicerates/Arachnids Myriapods Crustaceans Hexapods/Insects Platyhelminths Turbellaria Trematoda Monogenea Cestoda Bryozoans Phylactolaemata Stenolaemata Gymnolaemata Annelids Polychaetes Clitellata Echiura Molluscs Gastropods Cephalopods Bivalves Chitons Tusk shells Phyla with ≥5000 extant species bolded See also Diploblasts Monoblastozoa (nomen dubium) ¹Platyzoa ²Porifera (sponges) v t e Living things in culture Academic disciplines Anthrozoology Ethnobiology Ethnobotany Ethnoecology Ethnoentomology Ethnoherpetology Ethnoichthyology Ethnolichenology Ethnomycology Ethnoornithology Ethnoprimatology Ethnozoology Groups Animals Arthropods Insects Topics Beekeeping Entomophagy Flea circus Insects in art Insects in literature Insects in medicine Insects in music Insects in mythology Insects in religion Sericulture Types Ant Bee Mythology Beetle Beetlewing Butterfly Cicada Cricket Dragonfly Flea Flea circus Fly Grasshopper Ladybird Louse Praying mantis Scarab Termite Wasp Woodworm Other Arthropods in film Crab Lobster Scorpion Spider Arachnophobia Tarantella Tick Molluscs Conch (instrument) Conchology Pearl Scallop Seashell Sea silk Shell money Shipworm Tyrian purple List of edible molluscs Octopus Pitar dione Vertebrates Amphibians Frog Salamander Toad Toadstone Birds Aviculture Birdwatching Bird conservation Birds in culture Cockfighting Falconry Game bird Pigeon racing Poultry Archaeopteryx Barnacle goose Eagle Fish Fishing History Fish farming Fishkeeping Recreational fishing Mammals Topics Animal husbandry Fur farming Hunting In sport In professional wrestling Laboratory animal Livestock Pack animal Working animal Types Bat Bear Baiting Hunting Teddy bear Cattle Deer Elephant Fox Horse Riding Worship Leopard Lion Primate Gorilla Gorilla suit Monkey Orangutan Seal hunting Sheep Whale Tay Whale Whaling Whale watching Wolf Werewolf Reptiles Crocodile Attacks Farming Crocodile tears Dinosaur Dragon Lizard Snake Caduceus In the Bible Rod of Asclepius Snakebite Snake charming Symbolism Worship Turtle Bixi World Turtle Other phyla Coral Jellyfish Starfish Other Aesop's Fables Animal epithet Animal husbandry In heraldry Lists of legendary creatures Zodiac Plants Agriculture History Botanical illustration Floral design Ikebana Gardening Herbalism Fictional plants Magical plants Mandrake Medicinal plants Pharmacognosy Plant epithet Sacred grove In India Sacred plants Bodhi Tree Lime tree Sacred lotus Sacred herb In mythology Barnacle tree Fig Trees Yggdrasil Fungi Medicinal fungi Amanita muscaria Edible mushroom Agaricus bisporus Psilocybin mushroom Microbes Biological warfare Fermentation In food processing List of microbes Microbial art Microbes and Man Pathogen Protein production Bacteria Economic importance Tuberculosis Protist Virus Yeast Bread Beer Wine Related Anthropomorphism Legendary creature Lists of fictional species Parasitoid Template:Insects in culture Animals portal Biology portal Taxon identifiers Wd: Q25326 ADW: Mollusca EoL: 2195 Fauna Europaea: 11368 Fossilworks: 7805 ITIS: 69458 NCBI: 6447 WoRMS: 51 Authority control GND: 4189417-0 NDL: 00568039 Retrieved from "" Categories: MolluscsAnimal phylaExtant Cambrian first appearancesControversial taxaHidden categories: Wikipedia articles incorporating a citation from the 1911 Encyclopaedia Britannica with Wikisource referenceCS1 maint: Date and yearCS1 maint: Unfit urlPages using web citations with no URLCS1 maint: BOT: original-url status unknownAll articles with dead external linksArticles with dead external links from May 2017CS1 maint: Explicit use of et al.Good articlesArticles using diversity taxoboxArticles with 'species' microformatsArticles with dead external links from April 2017Wikipedia articles with GND identifiersArticles containing video clips

Navigation menu Personal tools Not logged inTalkContributionsCreate accountLog in Namespaces ArticleTalk Variants Views ReadEditView history More Search Navigation Main pageContentsFeatured contentCurrent eventsRandom articleDonate to WikipediaWikipedia store Interaction HelpAbout WikipediaCommunity portalRecent changesContact page Tools What links hereRelated changesUpload fileSpecial pagesPermanent linkPage informationWikidata itemCite this page Print/export Create a bookDownload as PDFPrintable version In other projects Wikimedia CommonsWikispeciesWikibooks Languages AfrikaansAlemannischالعربيةAragonésAsturianuAzərbaycancaتۆرکجهবাংলাBân-lâm-gúБашҡортсаБеларускаяБеларуская (тарашкевіца)‎БългарскиBosanskiBrezhonegБуряадCatalàCebuanoČeštinaCymraegDanskDeutschDiné bizaadEestiΕλληνικάEspañolEsperantoEuskaraفارسیFøroysktFrançaisGaeilgeGalegoગુજરાતી한국어Հայերենहिन्दीHornjoserbsceHrvatskiIdoBahasa IndonesiaInterlinguaÍslenskaItalianoעבריתBasa JawaქართულიҚазақшаKinyarwandaKiswahiliКыргызчаLatinaLatviešuLëtzebuergeschLietuviųLimburgsMagyarМакедонскиമലയാളംBahasa MelayuMìng-dĕ̤ng-ngṳ̄Nederlands日本語NapulitanoNordfriiskNorskNorsk nynorskOccitanଓଡ଼ିଆOʻzbekcha/ўзбекчаਪੰਜਾਬੀپنجابیPlattdüütschPolskiPortuguêsRomânăRuna SimiРусскийScotsSimple EnglishSlovenčinaSlovenščinaСрпски / srpskiSrpskohrvatski / српскохрватскиSuomiSvenskaTagalogதமிழ்TarandíneతెలుగుTetunไทยТоҷикӣTürkçeУкраїнськаVènetoTiếng ViệtWest-VlamsWinarayייִדיש粵語中文 Edit links This page was last edited on 12 January 2018, at 08:07. Text is available under the Creative Commons Attribution-ShareAlike License; additional terms may apply. By using this site, you agree to the Terms of Use and Privacy Policy. Wikipedia® is a registered trademark of the Wikimedia Foundation, Inc., a non-profit organization. Privacy policy About Wikipedia Disclaimers Contact Wikipedia Developers Cookie statement Mobile view (window.RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgPageParseReport":{"limitreport":{"cputime":"1.664","walltime":"1.910","ppvisitednodes":{"value":25194,"limit":1000000},"ppgeneratednodes":{"value":0,"limit":1500000},"postexpandincludesize":{"value":777967,"limit":2097152},"templateargumentsize":{"value":77168,"limit":2097152},"expansiondepth":{"value":26,"limit":40},"expensivefunctioncount":{"value":10,"limit":500},"entityaccesscount":{"value":8,"limit":400},"timingprofile":["100.00% 1634.775 1 -total"," 35.85% 586.073 2 Template:Reflist"," 26.22% 428.595 1 Template:Automatic_Taxobox"," 25.62% 418.792 1 Template:Taxobox/core"," 19.52% 319.031 36 Template:Navbox"," 18.04% 294.937 58 Template:Cite_journal"," 11.84% 193.485 23 Template:Clade"," 9.42% 154.027 43 Template:Period_start"," 8.06% 131.699 1 Template:Fossil_range"," 7.98% 130.509 1 Template:Taxobox/taxonomy"]},"scribunto":{"limitreport-timeusage":{"value":"0.854","limit":"10.000"},"limitreport-memusage":{"value":10555419,"limit":52428800}},"cachereport":{"origin":"mw1265","timestamp":"20180116151427","ttl":1900800,"transientcontent":false}}});});(window.RLQ=window.RLQ||[]).push(function(){mw.config.set({"wgBackendResponseTime":95,"wgHostname":"mw1251"});});

Mollusca - Photos and All Basic Informations

Mollusca More Links

This Is A Good Article. Follow The Link For More Information.PrecambrianCambrianOrdovicianSilurianDevonianCarboniferousPermianTriassicJurassicCretaceousPaleogeneNeogeneTonicella LineataPolyplacophoranTaxonomy (biology)EAnimalLophotrochozoaCarl Linnaeus10th Edition Of Systema NaturaeBiodiversityEnlargeCornu AspersumLand SnailPhylumInvertebrateHelp:IPA/EnglishExtant TaxonSpeciesMarine BiologyOrganismFreshwater MolluscTerrestrial MolluscsHabitatAnatomicalTaxonomicClass (biology)ExtinctCephalopodSquidCuttlefishOctopusNeurobiologyInvertebrateGiant SquidColossal SquidGastropodSnailSlugMantle (mollusc)ExcretionRadulaNervous SystemLimpetGastropod ShellProteinChitinCalcium CarbonateCoelomateCoelomHemocoelBloodCirculatory SystemOpen Circulatory SystemMucusCiliaVentral Nerve CordBivalveBrainEsophagusEyesReproductive SystemExternal FertilizationEgg (biology)TrochophoreLarvaeVeligerCephalopodsCambrianMyrWikipedia:Link RotLophotrochozoaFossilAnatomically Modern HumanPearlMother Of PearlTyrian PurpleSea SilkMoneyBlue-ringed OctopusOctopus ApollyonInflammationCone ShellNeurologicalSchistosomiasisEcosystemLatinAristotleCuttlefishMalacologyBrachiopodBryozoaTunicateMantle (mollusc)Respiration (physiology)ExcretionCalcareousSynapomorphyDorsoventralMantle (mollusc)SecreteCalcareousMantle CavityAnusGenitalVentral Nerve CordClass (biology)AplacophoraPolyplacophoraMonoplacophoraGastropodaCephalopodaBivalviaScaphopodaRadulaChitinNeomeniomorphaCecumMetanephridiaOdontophoreEnlargeGastropodSnailSlugCowryFreshwater SnailTerrestrial MolluscsArthropodChordatePhylumSnailSlugGastropodClamBivalveSquidCephalopodAbyssal ZoneFreshwater MolluscsEcosystemTropicsTemperateLatitudeCephalopodSquidCuttlefishOctopusNeurobiologyGiant SquidInvertebrateColossal SquidFreshwater MolluscTerrestrial MolluscsIUCN Red ListMollusc ShellEnlargeMonoplacophoraLimpetGastropod ShellHabitatAnusOsphradiumGillNephridiaGonadMollusc ShellNudibranchChitinConchiolinProteinCalcium CarbonatePeriostracumCobcrephoraAragoniteCalcitePeriostracumCalciteNacreEnlargeNatica ChemnitziCerithiumGulf Of CaliforniaPuerto PeñascoStatocystMucusLimpetCirculatory SystemOpen Circulatory SystemCoelomateCoelomHeartHemocoelBloodCoelomHydrostatic SkeletonRespiratory PigmentHemocyaninOxygenAtrium (heart)Ventricle (heart)AortaArteryExcretory SystemUrineNephridiaSedimentEnlargeRadulaRadulaOdontophoreIntracellular DigestionRadulaeBacteriaAlgaeOdontophoreGlandMucusCiliaFecesCecumDetritusEnlargeVentral Nerve CordVisceraCommissureEsophagusCircumesophageal Nerve RingScallopsReproductive System Of GastropodsReproductive System Of CephalopodsTrochophore Larva[36]File:Trochophore Larva 01.pngExternal FertilizationTrochophoreVeligerGonadCoelomOvumSpermExternal FertilizationInternal FertilizationHermaphroditeLarvaTrochophorePlanktonMesodermVeligerMetamorphosisFilter FeedersXenophyophoreDoridNudibranchsVetigastropodaList Of Mollusc OrdersClass (biology)CaudofoveataSolenogastersAplacophoraCaudofoveataSolenogastresPolyplacophoraMonoplacophoraGastropodaSnailSlugAbaloneLimpetConchNudibranchSea HareSea ButterflyCephalopodSquidOctopusCuttlefishNautilusSpirulaBivalviaClamOysterScallopGeoduckMusselScaphopodaRostroconchiaExtinctHelcionelloidaLatouchellaTentaculitaTentaculitesWiwaxiaEvolution Of MolluscsEvolution Of CephalopodsEnlargeLove DartMonachoides VicinusSexual SelectionGastropodAldanellaCephalopodPlectronocerasNectocarisBivalvePojetaiaFordillaCambrianLophotrochozoaFossilEdiacaranCambrianKimberellaBilateriaWiwaxiaRadulaPolychaeteMicrofossilMyrEnlargeHelcionellidYochelcionellaMolluscEnlargeGastropodHelcionellidSiberiaTrilobiteLimpetGastropodTorsion (gastropod)Septa And Siphuncle In Nautiloid ShellSiphuncleFile:Nautiloid Septa N Siphuncle 01.pngSiphuncleNautiloidVolborthellaSilicon DioxideSeptum (marine Biology)NautilusPlectronocerasSiphuncleAmmoniteNautiloidCretaceousColeoideaSquidOctopusCuttlefishFordillaPojetaiaBivalveOrdovicianMyrRudistReefCretaceous–Paleogene Extinction EventHyolithaOperculum (gastropod)PhylumLophotrochozoaBrachiopodBivalveMonoplacophoraGastropodSnailSlugLimpetSea HareCephalopodNautiloidAmmonitesSquidScaphopodAplacophoraPolyplacophoraHalwaxiidWiwaxiaHalkieriaOrthrozanclusOdontogriphusAnnelidPhylogenyKimberellaHalwaxiidLophotrochozoaTrochophoreLophophorataLophophoreAnnelidPhylumMetamerism (biology)AnnelidConchiferaAplacophoraEmbryologyCuticleEngrailed (gene)SkeletonAragoniteAculiferaSolenogastresCaudofoveataPolyplacophoraConchiferaMonoplacophoraBivalveScaphopodGastropodCephalopodAculiferaSolenogastresCaudofoveataTestariaPolyplacophoraMonoplacophoraBivalveScaphopodGastropodCephalopodTestariaMorphology (biology)Molecular PhylogeneticsMonophyleticScaphopodaBivalviaPolyphyleticMolluscs In CulturePearlMother Of PearlTyrian PurpleSea SilkCurrencySeashellList Of Edible MolluscsClamMusselOctopusSquidWhelkOystersScallopsSeafoodToxinEnlargePearl OysterIndonesiaBivalveGastropodNacrePearl OysterPinctadaTropicalSubtropicalPacific OceanMantle (mollusc)PearlsMusselEndangeredEnlargeByzantineJustinian ITyrian PurpleSocial StatusTyrian PurpleMurexBefore ChristTheopompusCreteMinoansTyriansSea SilkTextileByssusPinna NobilisProcopiusCommon EraArmeniaChlamysCowrieMoneyShell MoneySocial StatusCommodityInflationBioindicatorsPotamopyrgus AntipodarumEnlargeBlue-ringed OctopusJellyfishBlue-ringed OctopusOctopus ApollyonInflammationEast Pacific Red OctopusNecrosisEnlargeCone ShellNeurologyCone SnailToxinNeurologyMoleculeEnlargeVesicle (dermatology)SchistosomaCenters For Disease Control And PreventionSchistosomiasisSchistosomaHost (biology)SlugsEcosystemAchatina FulicaIndian OceanPacific OceanWest IndiesEuglandina RoseaAchatina FulicaInternational Standard Book NumberSpecial:BookSources/978-0-642-56860-1International Standard Book NumberSpecial:BookSources/978-0-642-56861-8Encyclopædia Britannica Eleventh EditionBibcodeDigital Object IdentifierPubMed CentralPubMed IdentifierInternational Standard Book NumberSpecial:BookSources/0-19-854055-8International Standard Book NumberSpecial:BookSources/0-87893-097-3International Standard Book NumberSpecial:BookSources/978-0-520-25092-5Digital Object IdentifierJSTORDigital Object IdentifierInternational Standard Book NumberSpecial:BookSources/0716700220Digital Object IdentifierPubMed CentralPubMed IdentifierBioScienceDigital Object IdentifierCategory:CS1 Maint: Date And YearCategory:CS1 Maint: Unfit UrlInternational Standard Book NumberSpecial:BookSources/0-19-551368-1BibcodeDigital Object IdentifierPubMed IdentifierDigital Object IdentifierDigital Object IdentifierDigital Object IdentifierHandle SystemInternational Standard Book NumberSpecial:BookSources/0-12-728702-7Digital Object IdentifierPubMed IdentifierInternational Standard Book NumberSpecial:BookSources/0030259827Digital Object IdentifierDigital Object IdentifierDigital Object IdentifierDigital Object IdentifierHelp:CS1 ErrorsInternational Standard Book NumberSpecial:BookSources/0-632-05238-4BibcodeDigital Object IdentifierJSTORPubMed IdentifierBibcodeDigital Object IdentifierPubMed IdentifierBibcodeDigital Object IdentifierBibcodeDigital Object IdentifierDigital Object IdentifierPubMed IdentifierDigital Object IdentifierPubMed CentralPubMed IdentifierBibcodeDigital Object IdentifierPubMed IdentifierDigital Object IdentifierDigital Object IdentifierJSTORPubMed IdentifierBibcodeDigital Object IdentifierInternational Standard Book NumberSpecial:BookSources/978-1-86239-233-5Digital Object IdentifierBibcodeDigital Object IdentifierDigital Object IdentifierJSTORCategory:CS1 Maint: Date And YearInternational Standard Book NumberSpecial:BookSources/0-520-25092-3Category:CS1 Maint: BOT: Original-url Status UnknownInternational Standard Book NumberSpecial:BookSources/0-486-29371-8BibcodeDigital Object IdentifierPubMed IdentifierDigital Object IdentifierCategory:CS1 Maint: Date And YearDigital Object IdentifierDigital Object IdentifierDigital Object IdentifierBibcodeDigital Object IdentifierPubMed IdentifierDigital Object IdentifierDigital Object IdentifierPubMed CentralPubMed IdentifierCladistics (journal)Digital Object IdentifierDigital Object IdentifierPubMed IdentifierDigital Object IdentifierPubMed IdentifierDigital Object IdentifierPubMed IdentifierDigital Object IdentifierPubMed IdentifierDigital Object IdentifierPubMed IdentifierDigital Object IdentifierPubMed CentralPubMed IdentifierDerek BriggsDigital Object IdentifierPubMed CentralPubMed IdentifierDigital Object IdentifierJSTORInternational Standard Book NumberSpecial:BookSources/0-443-05627-7FAODigital Object IdentifierCategory:CS1 Maint: Date And YearBefore ChristTheopompusInternational Standard Book NumberSpecial:BookSources/0-674-99380-2Digital Object IdentifierDigital Object IdentifierJSTORDigital Object IdentifierWikipedia:Link RotInternational Standard Book NumberSpecial:BookSources/0521541107Category:CS1 Maint: Explicit Use Of Et Al.International Standard Book NumberSpecial:BookSources/0-86840-279-6Digital Object IdentifierPubMed IdentifierMelbourneDigital Object IdentifierPubMed IdentifierCarter CenterInternational Standard Book NumberSpecial:BookSources/0-7484-0026-5International Standard Book NumberSpecial:BookSources/0-85199-320-6Digital Object IdentifierInternational Standard Book NumberSpecial:BookSources/0-03-025982-7International Standard Book NumberSpecial:BookSources/1581129300Wikispecies:MolluscaEncyclopedia Of LifeTemplate:AnimaliaTemplate Talk:AnimaliaAnimalPhylumDomain (biology)Template:Archaea ClassificationTemplate:Bacteria ClassificationTemplate:EukaryotaKingdom (biology)Template:Plant ClassificationTemplate:HacrobiaTemplate:HeterokontTemplate:AlveolataTemplate:RhizariaTemplate:ExcavataTemplate:AmoebozoaTemplate:Opisthokont ProtistsTemplate:AnimaliaTemplate:Fungi ClassificationAnimalHexactinellidDemospongeCalcareous SpongeHomoscleromorphaDiploblastEumetazoaCtenophoraParaHoxozoaPlacozoaPlanulozoaCnidariaBilateriaTriploblastsBilateriaXenacoelomorphaXenoturbellaAcoelomorphaAcoelaNemertodermatidaNephrozoaDeuterostomeChordateLanceletTunicateCraniateVertebrateAmbulacrariaEchinodermHemichordateAcorn WormPterobranchiaProtostomeEcdysozoaScalidophoraKinorhynchaPriapulidaNematoidaNematodeNematomorphaLoriciferaPanarthropodaOnychophoraTactopodaArthropodTardigradeSpiraliaGnathifera (clade)ChaetognathaGnathostomulidLimnognathiaSyndermataRotiferAcanthocephalaPlatytrochozoaMesozoaOrthonectidaDicyemidaRouphozoaFlatwormGastrotrichLophotrochozoaSymbionAnnelidNemerteaLophophorataBryozoaEntoproctaBryozoaBrachiozoaBrachiopodPhoronidSpongeCalcareous SpongeHexactinellidDemospongeHomoscleromorphaCnidariaAnthozoaMedusozoaMyxozoaVertebrateAgnathaChondrichthyesOsteichthyesAmphibianReptileBirdMammalEchinodermCrinozoaAsterozoaEchinozoaNematodeChromadoreaEnopleaSecernenteaArthropodChelicerataMyriapodaCrustaceanHexapodaFlatwormTurbellariaTrematodaMonogeneaCestodaBryozoaPhylactolaemataStenolaemataGymnolaemataAnnelidPolychaeteClitellataEchiuraGastropodaCephalopodBivalviaChitonTusk ShellDiploblastSalinellaNomen DubiumPlatyzoaSpongeTemplate:Living Things In CultureTemplate Talk:Living Things In CultureLiving Things In CultureAnthrozoologyEthnobiologyEthnobotanyEthnoecologyEthnoentomologyEthnoherpetologyEthnoichthyologyEthnolichenologyEthnomycologyEthnoornithologyEthnoprimatologyEthnozoologyAnimals In CultureArthropods In CultureInsects In CultureBeekeepingEntomophagyFlea CircusInsects In ArtInsects In LiteratureInsects In MedicineInsects In MusicInsects In MythologyInsects In ReligionSericultureAntBeeBee (mythology)BeetleBeetlewingButterflyCicada (mythology)Cricket (insect)DragonflyFleaFlea CircusFlyGrasshopperLadybirdLousePraying MantisScarab (artifact)TermiteWaspWoodwormArthropods In FilmCrabLobsterScorpionCultural Depictions Of SpidersArachnophobiaTarantellaTickMolluscs In CultureConch (instrument)ConchologyPearlScallopSeashellSea SilkShell MoneyTeredo NavalisTyrian PurpleList Of Edible MolluscsOctopusPitar DioneAmphibians In CultureFrogs In CultureSalamanders In Folklore And LegendToadToadstoneBirds In CultureAvicultureBirdwatchingBird ConservationBirds In CultureCockfightFalconryGame (hunting)Pigeon RacingPoultryArchaeopteryxBarnacle GooseEagleFish In CultureFishingHistory Of FishingFish FarmingFishkeepingRecreational FishingMammals In CultureAnimal HusbandryFur FarmingHuntingAnimals In SportAnimals In Professional WrestlingLaboratory AnimalLivestockPack AnimalWorking AnimalBatBearBear-baitingBear HuntingTeddy BearCattleDeer In MythologyCultural Depictions Of ElephantsFoxes In Popular CultureHorseEquestrianismHorse WorshipLeopards In CultureCultural Depictions Of LionsPrimateGorillas In Popular CultureGorilla SuitMonkeyOrangutans In Popular CulturePinnipedSeal HuntingSheepWhaleTay WhaleWhalingWhale WatchingWolves In Folklore, Religion And MythologyWerewolfReptiles In CultureCrocodileCrocodile AttackCrocodile FarmCrocodile TearsCultural Depictions Of DinosaursDragonLizardSnakeCaduceusSerpents In The BibleRod Of AsclepiusSnakebiteSnake CharmingSerpent (symbolism)Snake WorshipTurtleBixi (mythology)World TurtleCoralJellyfishStarfishAesop's FablesAnimal EpithetAnimal HusbandryCharge (heraldry)Lists Of Legendary CreaturesZodiacPlants In CultureAgricultureHistory Of AgricultureBotanical IllustrationFloral DesignIkebanaGardeningHerbalismList Of Fictional PlantsMandrakeMedicinal PlantsPharmacognosyPlant EpithetSacred GroveSacred Groves Of IndiaBodhi TreeLime Tree In CultureNelumbo NuciferaSacred HerbBarnacle GooseCommon FigTrees In MythologyYggdrasilFungi In Human CultureMedicinal FungiAmanita MuscariaEdible MushroomAgaricus BisporusPsilocybin MushroomMicrobes In Human CultureBiological WarfareFermentationFermentation In Food ProcessingList Of Microorganisms Used In Food And Beverage PreparationMicrobial ArtMicrobes And ManPathogenProtein ProductionBacteriaEconomic Importance Of BacteriaTuberculosis In Human CultureProtistVirusYeastBread In CultureBeerWineAnthropomorphismLegendary CreatureLists Of Fictional SpeciesParasitoidTemplate:Insects In CulturePortal:AnimalsPortal:BiologyHelp:Taxon IdentifiersWikidataAnimal Diversity WebEncyclopedia Of LifeFauna EuropaeaFossilworksIntegrated Taxonomic Information SystemNational Center For Biotechnology InformationWorld Register Of Marine SpeciesHelp:Authority ControlIntegrated Authority FileNational Diet LibraryHelp:CategoryCategory:MolluscsCategory:Animal PhylaCategory:Extant Cambrian First AppearancesCategory:Controversial TaxaCategory:Wikipedia Articles Incorporating A Citation From The 1911 Encyclopaedia Britannica With Wikisource ReferenceCategory:CS1 Maint: Date And YearCategory:CS1 Maint: Unfit UrlCategory:Pages Using Web Citations With No URLCategory:CS1 Maint: BOT: Original-url Status UnknownCategory:All Articles With Dead External LinksCategory:Articles With Dead External Links From May 2017Category:CS1 Maint: Explicit Use Of Et Al.Category:Good ArticlesCategory:Articles Using Diversity TaxoboxCategory:Articles With 'species' MicroformatsCategory:Articles With Dead External Links From April 2017Category:Wikipedia Articles With GND IdentifiersCategory:Articles Containing Video ClipsDiscussion About Edits From This IP Address [n]A List Of Edits Made From This IP Address [y]View The Content Page [c]Discussion About The Content Page [t]Edit This Page [e]Visit The Main Page [z]Guides To Browsing WikipediaFeatured Content – The Best Of WikipediaFind Background Information On Current EventsLoad A Random Article [x]Guidance On How To Use And Edit WikipediaFind Out About WikipediaAbout The Project, What You Can Do, Where To Find ThingsA List Of Recent Changes In The Wiki [r]List Of All English Wikipedia Pages Containing Links To This Page [j]Recent Changes In Pages Linked From This Page [k]Upload Files [u]A List Of All Special Pages [q]Wikipedia:AboutWikipedia:General Disclaimer