Contents 1 Structure 1.1 Microanatomy 1.2 Detrusor muscle 1.3 Blood- and lymph-supply 1.4 Nerve supply 1.5 Development 2 Function 3 Clinical significance 3.1 Inflammation and infection 3.2 Incontinence and retention 3.3 Cancer 3.4 Investigation 4 Other animals 4.1 Reptiles 4.2 Amphibians 4.3 Fish 4.4 Mammals 4.5 Birds 5 Additional images 6 See also 7 References 8 External links

Structure[edit] Further information: Urination § Anatomy of the bladder and outlet Male and female urinary bladders in lateral cross-section Bladder location and associated structures in the male In humans the bladder is a hollow muscular organ situated at the base of the pelvis. Urine collects in the bladder, fed from the two ureters that connect the bladder with the kidneys. Urine leaves the bladder via the urethra, a single muscular tube which ends in an opening – the urinary meatus, where it exits the body. Anatomists divide the bladder into:.[3] a broad fundus a body an apex a neck The apex (prev.vertex) is directed forward toward the upper part of the pubic symphysis, and from there the median umbilical ligament continues upward on the back of the anterior abdominal wall to the umbilicus. The peritoneum is carried by it from the apex on to the abdominal wall to form the middle umbilical fold. The neck of the bladder is the area at the base of the trigone that surrounds the internal urethral orifice that leads to the urethra.[3] In males the neck of the urinary bladder is adjacent to the prostate gland. The three openings, two ureteric orifices, and the internal urethral orifice mark the triangular area called the trigone of the bladder. These openings have mucosal flaps in front of them that act as valves in preventing the backflow of urine into the ureters,[4] known as vesicoureteral reflux. Between the two ureteric openings is a raised area of tissue called the interureteric crest.[3] This makes the upper boundary of the trigone. The trigone is a smooth-muscle area that forms the floor of the bladder above the urethra.[5] It is an area of smooth tissue for the easy flow of urine into and from this part of the bladder - in contrast to the irregular surface formed by the rugae. In men, the prostate gland lies outside the opening for the urethra. The middle lobe of the prostate causes an elevation in the mucous membrane behind the internal urethral orifice called the uvula of urinary bladder. The uvula can enlarge when the prostate becomes enlarged. The bladder is situated below the peritoneal cavity near the pelvic floor and behind the pubic symphysis. In men, it lies in front of the rectum, separated by the recto-vesical pouch, and is supported by fibres of the levator ani and of the prostate gland. In women, it lies in front of the uterus, separated by the vesico-uterine pouch, and is supported by the elevator ani and the upper part of the vagina. The wall of the urinary bladder is normally 3–5 mm thick.[6] When well distended, the wall is normally less than 3 mm.[6] The inner walls have a series of ridges, thick mucosal folds known as rugae that allow for the expansion of the bladder. The Latin phrase for "urinary bladder" is vesica urinaria, and the term vesical or prefix vesico - appear in connection with associated structures such as vesical veins. The modern Latin word for "bladder" - cystis - appears in associated terms such as cystitis (inflammation of the bladder). Microanatomy[edit] The outside of the bladder is protected by a serous membrane.[7] The bladder wall itself is smooth muscle.[7] The inner side of the bladder is lined with a mucosal membrane consisting of a surface glycocalyx that protects the cells beneath it from urine, the urothelium (a form of transitional epithelium), a basement membrane, and the lamina propria.[7][8] The mucosal lining also offers a urothelial barrier against the passing of infections.[9] Vertical section of bladder wall. Layers of the urinary bladder wall and cross section of the detrusor muscle. Anatomy of the male bladder, showing transitional epithelium and part of the wall in a histological cut-out. Detrusor muscle[edit] The detrusor muscle is a layer of the urinary bladder wall made of smooth muscle fibers arranged in spiral, longitudinal, and circular bundles. Stretch receptors in the bladder signal the parasympathetic nervous system to stimulate the muscarinic receptors in the detrusor to contract the muscle when the bladder is extended.[10] This encourages the bladder to expel urine through the urethra. The main receptor activated is the M3 receptor, although M2 receptors are also involved and whilst outnumbering the M3 receptors they are not so responsive.[11] The main relaxant pathway is via the adenylyl cyclase cAMP pathway, activated by β2 adrenergic receptors.[12] The detrusor muscle is unusually able to change its length. It can also contract for a long time whilst voiding, and it stays relaxed whilst the bladder is filling.[12] Blood- and lymph-supply[edit] The bladder is supplied by the vesical arteries and drained by the vesical veins.[13] The superior vesical artery supplies blood to the upper part of the bladder, and the inferior vesical artery, supplies the lower part and its just in male (insted of this in female is vaginal artery( . Both are branches of the internal iliac arteries.[13] The uterine arteries and vaginal arteries also supply blood in females.[13] Venous drainage of blood supplying the bladder begins in a network of small vessels on the outer and back surfaces of the bladder. These coalesce and pass backwards along the lateral ligaments of the bladder into the internal iliac veins.[13] The lymph drained from the bladder begins in a series of networks throughout the mucosal, muscular and serosal layers. These then form three sets of vessels: one set near the trigone draining the bottom of the bladder; one set draining the top of the bladder; and another set draining the outer undersurface of the bladder. The majority of these vessels drain into the external iliac lymph nodes.[13] Nerve supply[edit] The bladder receives motor innervation from both sympathetic fibers, most of which arise from the superior and inferior hypogastric plexuses and nerves, and from parasympathetic fibers, which come from the pelvic splanchnic nerves.[14] Sensation from the bladder is transmitted to the central nervous system (CNS) via general visceral afferent fibers (GVA). GVA fibers on the superior surface follow the course of the sympathetic efferent nerves back to the CNS, while GVA fibers on the inferior portion of the bladder follow the course of the parasympathetic efferents.[14] For the urine to exit the bladder, both the autonomically controlled internal sphincter (in the male) and the voluntarily controlled external sphincter must be opened. Problems with these muscles can lead to incontinence.[15] Development[edit] The human urinary bladder derives from the urogenital sinus, and it is initially continuous with the allantois. The upper and lower parts of the bladder develop separately and join together around the middle part of development.[5] At this time the ureters move from the mesonephric ducts to the trigone.[5] In males, the base of the bladder lies between the rectum and the pubic symphysis. It is superior to the prostate, and separated from the rectum by the recto-vesical pouch. In females, the bladder sits inferior to the uterus and anterior to the vagina; thus its maximum capacity is lower than in males. It is separated from the uterus by the vesico-uterine pouch. In infants and young children the urinary bladder is in the abdomen even when empty.[16]

Function[edit] Main article: Urination Urine, excreted by the kidneys, collects in the bladder before disposal by urination (micturition). The urinary bladder usually holds 300-350 ml of urine. As urine accumulates, the rugae flatten and the wall of the bladder thins as it stretches, allowing the bladder to store larger amounts of urine without a significant rise in internal pressure.[17] Urination is controlled by the pontine micturition center in the brainstem.[18]

Clinical significance[edit] See also: Urinary bladder disease Inflammation and infection[edit] Calcifiications on bladder wall caused by urinary schistosomiasis Bacteriuria is the presence of bacteria in the urine which can indicate a urinary tract infection such as cystitis. Interstitial cystitis Incontinence and retention[edit] Frequent urination can be due to excessive urine production, small bladder capacity, irritability or incomplete emptying. Males with an enlarged prostate urinate more frequently. One definition of an overactive bladder is when a person urinates more than eight times per day.[19] An overactive bladder can often cause urinary incontinence. Though both urinary frequency and volumes have been shown to have a circadian rhythm, meaning day and night cycles,[20] it is not entirely clear how these are disturbed in the overactive bladder. Urodynamic testing can help to explain the symptoms. An underactive bladder is the condition where there is a difficulty in passing urine and is the main symptom of a neurogenic bladder. Frequent urination at night may indicate the presence of bladder stones. Disorders of or related to the bladder include: Bladder exstrophy Bladder sphincter dyssynergia, a condition in which the sufferer cannot coordinate relaxation of the urethra sphincter with the contraction of the bladder muscles Paruresis Trigonitis Urinary retention Disorders of bladder function may be dealt with surgically, by re-directing the flow of urine or by replacement with an artificial urinary bladder. The volume of the bladder may be increased by bladder augmentation. An obstruction of the bladder neck may be severe enough to warrant surgery. Cancer[edit] Bladder cancer – the most common type is transitional cell carcinoma.[21] Investigation[edit] A diverticulum of the bladder Urodynamic testing can help to explain the symptoms.

Other animals[edit] Reptiles[edit] In all reptiles the urinogenital ducts and the anus both empty into an organ called a cloaca. In some reptiles, a midventral wall in the cloaca may open into a urinary bladder, but not all. It is present in all turtles and tortoises as well as most lizards, but is lacking in the monitor lizard, the legless lizards. It is absent in the snakes, alligators, and crocodiles.[22]:p. 474 Many turtles, tortoises, and lizards have proportionally very large bladders. Charles Darwin noted that the Galapagos tortoise had a bladder which could store up to 20% of its body weight.[23] Such adaptations are the result of environments such as remote islands and deserts where water is very scarce.[24]:143 Other desert-dwelling reptiles have large bladders that can store a long-term reservoir of water for up to several months and aid in osmoregulation.[25] Turtles have two or more accessory urinary bladders, located lateral to the neck of the urinary bladder and dorsal to the pubis, occupying a significant portion of their body cavity.[26] Their bladder is also usually bilobed with a left and right section. The right section is located under the liver, which prevents large stones from remaining in that side while the left section is more likely to have calculi.[27] Amphibians[edit] Most aquatic and semi-aquatic amphibians have a membranous skin which allows them to absorb water directly through it. Some semi-aquatic animals also have similarly permeable bladder membrane.[28] As as result, they tend to have high rates of urine production to offset this high water intake, and have urine which is low in dissolved salts. The urinary bladder assists such animals to retain salts. Some aquatic amphibian such as Xenopus do not reabsorb water, to prevent excessive water influx.[29] For land-dwelling amphibians, dehydration results in reduced urine output.[30]:p. 184 The amphibian bladder is usually highly distensible and among some land-dwelling species of frogs and salamanders may account for between 20% and 50% of their total body weight.[30]:p. 184 Fish[edit] Although the gills of most teleost fish assist with the elimination of ammonia from the body and although fish virtually by definition live in an environment surrounded by water, most still have a distinct bladder for storing waste fluid. The urinary bladder of teleosts is permeable to water, though this is less true for freshwater dwelling species than saltwater ones.[23]:p. 219 Most fish also have an organ called a swim-bladder which is unrelated to the urinary bladder except in its membranous nature. The loaches, pilchards, and herrings are among the few types of fish in which a urinary bladder is poorly developed. It is largest in those fish which lack an air bladder, and is situated in front of the oviducts and behind the rectum.[31] Mammals[edit] All mammals have a urinary bladder. This structure begins as an embryonic cloaca. In the vast majority, this eventually becomes differentiated into a dorsal part connected to the intestine and a ventral part which becomes associated with the urinogenital passage and urinary bladder. The only mammals in which this does not take place are the platypus and the spiny anteater both of which retain the cloaca into adulthood.[22] The mammalian bladder is an organ that regularly stores a hyperosmotic concentration of urine. It therefore is relatively impermeable and has multiple epithelial layers. The urinary bladder of the cetaceans (whales and dolphins) is proportionally smaller than that of land-dwelling mammals.[32] Birds[edit] In the whole class of Aves there is no urinary bladder per se. Although all birds have kidneys, the ureters open directly into a cloaca which serves as a reservoir for urine, fecal matter, and eggs.[33]

Additional images[edit] Urinary bladder (black butterfly-like shape) and hyperplastic prostate (BPH) visualized by medical ultrasound.

See also[edit] Alpha blocker Cystitis glandularis Ureterocele

References[edit] ^ Boron, Walter F.; Boulpaep, Emile L. (2016). Medical Physiology. 3: Elsevier Health Sciences. p. 738. ISBN 9781455733286. Retrieved 1 June 2016.  ^ Walker-Smith, John; Murch, Simon (1999). Cardozo, Linda, ed. Diseases of the Small Intestine in Childhood (4 ed.). CRC Press. p. 16. ISBN 9781901865059. Retrieved 1 June 2016.  ^ a b c Netter, Frank H. (2014). Atlas of Human Anatomy Including Student Consult Interactive Ancillaries and Guides (6th ed.). Philadelphia, Penn.: W B Saunders Co. pp. 346–348. ISBN 978-14557-0418-7.  ^ "SEER Training:Urinary Bladder".  ^ a b c Viana, R; et al. (October 2007). "The development of the bladder trigone, the center of the anti-reflux mechanism". Development. 134 (20): 3763–9. PMID 17881488. CS1 maint: Explicit use of et al. (link) ^ a b Page 12 in: Uday Patel (2010). Imaging and Urodynamics of the Lower Urinary Tract. Springer Science & Business Media. ISBN 9781848828360.  ^ a b c Fry, CH; Vahabi, B (October 2016). "The Role of the Mucosa in Normal and Abnormal Bladder Function". Basic & clinical pharmacology & toxicology. 119 Suppl 3: 57–62. doi:10.1111/bcpt.12626. PMID 27228303.  ^ Merrill, L; et al. (April 2016). "Receptors, channels, and signalling in the urothelial sensory system in the bladder". Nature reviews. Urology. 13 (4): 193–204. PMID 26926246. CS1 maint: Explicit use of et al. (link) ^ Janssen, DA (January 2013). "The distribution and function of chondroitin sulfate and other sulfated glycosaminoglycans in the human bladder and their contribution to the protective bladder barrier". The Journal of Urology. 189 (1): 336–42. PMID 23174248.  ^ Giglio, D; Tobin, G (2009). "Muscarinic receptor subtypes in the lower urinary tract". Pharmacology. 83 (5): 259–69. PMID 19295256.  ^ Uchiyama, T; Chess-Williams, R (December 2004). "Muscarinic receptor subtypes of the bladder and gastrointestinal tract". Journal of smooth muscle research = Nihon Heikatsukin Gakkai kikanshi. 40 (6): 237–47. PMID 15725706.  ^ a b Andersson, K.-E. (1 July 2004). "Urinary Bladder Contraction and Relaxation: Physiology and Pathophysiology". Physiological Reviews. 84 (3): 935–986. doi:10.1152/physrev.00038.2003.  ^ a b c d e Gray's 2008, p. 1249. ^ a b Moore, Keith; Anne Agur (2007). Essential Clinical Anatomy, Third Edition. Lippincott Williams & Wilkins. pp. 227–228. ISBN 0-7817-6274-X.  ^ "Urinary Incontinence - Causes". NHS. Retrieved 2013-08-21.  ^ Moore, Keith L.; Dalley, Arthur F (2006). Clinically Oriented Anatomy (5th ed.). Lippincott Williams & Wilkins.  ^ Marieb, Mallatt. "23". Human Anatomy (5th ed.). Pearson International. p. 700.  ^ Purves, Dale (2011). Neuroscience (5. ed.). Sunderland, Mass.: Sinauer. p. 471. ISBN 978-0-87893-695-3.  ^ "Overactive Bladder". Cornell Medical College. Retrieved 2013-08-21.  ^ Negoro, Hiromitsu (2012). "Involvement of urinary bladder Connexin43 and the circadian clock in coordination of diurnal micturition rhythm". doi:10.1038/ncomms1812.  ^ "Bladder Cancer". National Cancer Institute. Archived from the original on 17 July 2017.  ^ a b Herbert W. Rand (1950). The Chordates. Balkiston.  ^ a b P.J. Bentley (14 March 2013). Endocrines and Osmoregulation: A Comparative Account in Vertebrates. Springer Science & Business Media. ISBN 978-3-662-05014-9.  ^ Paré, Jean (January 11, 2006). "Reptile Basics: Clinical Anatomy 101" (PDF). Proceedings of the North American Veterinary Conference. 20: 1657–1660.  ^ Davis, Jon R.; DeNardo, Dale F. (2007-04-15). "The urinary bladder as a physiological reservoir that moderates dehydration in a large desert lizard, the Gila monster Heloderma suspectum". Journal of Experimental Biology. 210 (8): 1472–1480. doi:10.1242/jeb.003061. ISSN 0022-0949. PMID 17401130.  ^ Wyneken, Jeanette; Witherington, Dawn (February 2015). "Urogenital System" (PDF). Anatomy of Sea Turtles. 1: 153–165.  ^ Divers, Stephen J.; Mader, Douglas R. (2005). Reptile Medicine and Surgery. Amsterdam: Elsevier Health Sciences. pp. 481, 597. ISBN 9781416064770.  ^ Urakabe, Shigeharu; Shirai, Dairoku; Yuasa, Shigekazu; Kimura, Genjiro; Orita, Yoshimasa; Abe, Hiroshi. "Comparative study of the effects of different diuretics on the permeability properties of the toad bladder". Comparative Biochemistry and Physiology Part C: Comparative Pharmacology. 53 (2): 115–119. doi:10.1016/0306-4492(76)90063-0.  ^ Shibata, Yuki; Katayama, Izumi; Nakakura, Takashi; Ogushi, Yuji; Okada, Reiko; Tanaka, Shigeyasu; Suzuki, Masakazu. "Molecular and cellular characterization of urinary bladder-type aquaporin in Xenopus laevis". General and Comparative Endocrinology. 222: 11–19. doi:10.1016/j.ygcen.2014.09.001.  ^ a b Laurie J. Vitt; Janalee P. Caldwell (25 March 2013). Herpetology: An Introductory Biology of Amphibians and Reptiles. Academic. ISBN 978-0-12-386920-3.  ^ Owen, Richard (1843). Lectures on the comparative anatomy and physiology of the invertebrate animals. London: Longman, Brown, Green, and Longmans. pp. 283–284.  ^ John Hunter (26 March 2015). The Works of John Hunter, F.R.S. Cambridge University. p. 35. ISBN 978-1-108-07960-0.  ^ Charles Knight (1854). The English Cyclopaedia: A New Dictionary of Universal Knowledge. Bradbury and Evans. p. 136.  Books editor-in-chief, Susan Standring ; section editors, Neil R. Borley; et al. (2008). Gray's anatomy : the anatomical basis of clinical practice (40th ed.). London: Churchill Livingstone. ISBN 978-0-8089-2371-8. CS1 maint: Explicit use of et al. (link) CS1 maint: Multiple names: authors list (link) CS1 maint: Extra text: authors list (link) Britton, the editors Nicki R. Colledge, Brian R. Walker, Stuart H. Ralston ; illustated by Robert (2010). Davidson's principles and practice of medicine (21st ed.). Edinburgh: Churchill Livingstone/Elsevier. 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External links[edit] Wikimedia Commons has media related to Urinary bladder. Histology at KUMC epithel-epith09 "Urinary Bladder" Anatomy photo: Urinary/mammal/bladder/bladder1 - Comparative Organology at University of California, Davis — "Mammal, bladder (LM, Medium)" Virtual Slidebox at Univ. Iowa Slide 445 Anatomy photo:43:07-0100 at the SUNY Downstate Medical Center — "The Female Pelvis: The Urinary bladder" Anatomy photo:44:04-0103 at the SUNY Downstate Medical Center — "The Male Pelvis: The Urinary bladder" Bladder (ISSN 2327-2120) — An open-access journal on bladder biology and diseases. v t e Human systems and organs Musculoskeletal Skeletal system Bone Carpus Collar bone (clavicle) Thigh bone (femur) Fibula Humerus Mandible Metacarpus Metatarsus Ossicles Patella Phalanges Radius Skull Tarsus Tibia Ulna Rib Vertebra Pelvis Sternum Cartilage Joints Fibrous joint Cartilaginous joint Synovial joint Muscular system Muscle Tendon Diaphragm Circulatory system Cardiovascular system peripheral Artery Vein Lymphatic vessel Heart Lymphatic system primary Bone marrow Thymus secondary Spleen Lymph node CNS equivalent Glymphatic system Nervous system Brain Spinal cord Nerve Sensory system Ear Eye Integumentary system Skin Subcutaneous tissue Breast Mammary gland Immune system Myeloid Myeloid immune system Lymphoid Lymphoid immune system Respiratory system Upper Nose Nasopharynx Larynx Lower Trachea Bronchus Lung Digestive system Mouth Salivary gland Tongue upper GI Oropharynx Laryngopharynx Esophagus Stomach lower GI Small intestine Appendix Colon Rectum Anus accessory Liver Biliary tract Pancreas Urinary system Genitourinary system Kidney Ureter Bladder Urethra Reproductive system Female Uterus Vagina Vulva Ovary Placenta Male Scrotum Penis Prostate Testicle Seminal vesicle Endocrine system Pituitary Pineal Thyroid Parathyroid Adrenal Islets of Langerhans v t e Anatomy of the urinary system Kidneys Layers Fascia Capsule Cortex column Medulla sinus pyramids medullary interstitium Lobe Cortical lobule Medullary ray Nephron Circulation Arteries Renal artery segmental interlobar arcuate interlobular afferent Veins Renal vein Peritubular capillaries Vasa recta arcuate interlobar efferent Nephron Renal corpuscle Glomerulus Bowman's capsule Glomerular basement membrane Podocyte Filtration slits Mesangium Intraglomerular mesangial cell Renal tubule Proximal convoluted tubule Loop of Henle Descending Thin ascending Thick ascending Distal convoluted tubule Connecting tubule Collecting ducts Tubular fluid Renal papilla Minor calyx Major calyx Renal pelvis Juxtaglomerular apparatus Macula densa Juxtaglomerular cells Mesangium Extraglomerular mesangial cell Ureters Ureteropelvic junction Bladder Apex Uvula Neck Median umbilical ligament Muscular layer Trigone Detrusor Mucosa Submucosa Urethra Urethral sphincters External sphincter male female Internal sphincter Authority control GND: 4072323-9 NDL: 00560840 Retrieved from "" Categories: PelvisOrgans (anatomy)Urinary systemUrinary bladderHidden categories: CS1 maint: Explicit use of et al.Medicine infobox template using GraySubject or GrayPageMedicine infobox template using Dorlands parameterCS1 maint: Multiple names: authors listCS1 maint: Extra text: authors listWikipedia articles with GND identifiersUse dmy dates from April 2017

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CenterBrainstemUrinary Bladder DiseaseEnlargeSchistosomiasisBacteriuriaUrinary Tract InfectionCystitisInterstitial CystitisEnlarged ProstateOveractive BladderUrinary IncontinenceUrodynamic TestingUnderactive BladderNeurogenic BladderBladder StoneBladder ExstrophyBladder Sphincter DyssynergiaParuresisTrigonitisUrinary RetentionArtificial Urinary BladderBladder AugmentationUrinary Bladder Neck ObstructionBladder CancerTransitional Cell CarcinomaEnlargeDiverticulumUrodynamic TestingAnusCloacaMonitor LizardLegless LizardCharles DarwinGalapagos TortoiseOsmoregulationBladder Stone (animal)XenopusTeleostSwim-bladderLoachPilchardHerringOviductRectumPlatypusSpiny AnteaterCetaceanAvesUretersCloacaProstateMedical UltrasoundAlpha BlockerCystitis GlandularisUreteroceleInternational Standard Book NumberSpecial:BookSources/9781455733286International Standard Book NumberSpecial:BookSources/9781901865059International Standard Book NumberSpecial:BookSources/978-14557-0418-7PubMed IdentifierCategory:CS1 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ListCategory:CS1 Maint: Extra Text: Authors ListInternational Standard Book NumberSpecial:BookSources/978-0-7020-3085-7University Of KansasUniversity Of IowaInternational Standard Serial NumberTemplate:Human Systems And OrgansTemplate Talk:Human Systems And OrgansHuman BodyOrgan (anatomy)Human Musculoskeletal SystemHuman SkeletonBoneCarpal BonesClavicleFemurFibulaHumerusMandibleMetacarpal BonesMetatarsal BonesOssiclesPatellaPhalanx BoneRadius (bone)SkullTarsus (skeleton)TibiaUlnaRibVertebral ColumnPelvisSternumCartilageJointFibrous JointCartilaginous JointSynovial JointMuscular SystemMuscleTendonThoracic DiaphragmCirculatory SystemCirculatory SystemArteryVeinLymphatic VesselHeartLymphatic SystemBone MarrowThymusSpleenLymph NodeGlymphatic SystemNervous SystemHuman BrainSpinal CordNerveSensory SystemEarHuman EyeIntegumentary SystemHuman SkinSubcutaneous TissueBreastMammary GlandImmune SystemMyeloidImmune SystemLymphocyteImmune SystemRespiratory SystemRespiratory TractHuman 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