Contents 1 Botanical berries 1.1 Modified berries 2 Fruits not botanical berries 2.1 Drupes 2.2 Pomes 2.3 Aggregate fruits 2.4 Multiple fruits 2.5 Accessory fruits 3 Berry-like conifer seed cones 4 History of terminology 5 Evolution and phylogenetic significance 6 Uses 6.1 Culinary 6.2 Others 6.3 History 6.4 Commercial production 7 See also 8 Notes 9 References 10 External links


Botanical berries[edit] Diagram of a grape berry, showing the pericarp and its layers Coffee cherries (Coffea arabica) – described as drupes or berries In botanical language, a berry is a simple fruit having seeds and fleshy pulp (the pericarp) produced from the ovary of a single flower. The ovary can be inferior or superior. It is indehiscent, i.e. it does not have a special "line of weakness" along which it splits to release the seeds when ripe.[2] The pericarp is divided into three layers. The outer layer is called the "exocarp" or "epicarp"; the middle layer, the "mesocarp" or "sarcocarp"; the inner layer, the "endocarp". Botanists have not applied these terms consistently. Exocarp and endocarp may be restricted to more-or-less single-layered "skins", or may include tissues adjacent to them; thus on one view, the exocarp extends inwards to the layer of vascular bundles ("veins"). The inconsistency in usage has been described as "a source of confusion".[3] The nature of the endocarp distinguishes a berry from a drupe, which has a hardened or stony endocarp (see also below). The two kinds of fruit intergrade, depending on the state of the endocarp. Some sources have attempted to quantify the difference, e.g. requiring the endocarp to be less than 2 mm thick in a berry.[4] Examples of botanical berries include: Avocado contains a single large seed surrounded by an imperceptible endocarp.[5] Banana[6][7] Barberry (Berberis), Oregon-grape (Mahonia aquifolium) and mayapple (Podophyllum spp.) (Berberidaceae) Strawberry tree (Arbutus unedo) (not to be confused with the strawberry (Fragaria), which is an accessory fruit), bearberry (Arctostaphylos spp.), bilberry, blueberry, cranberry, lingonberry/cowberry (Vaccinium vitis-idaea), crowberry (Empetrum spp.) (family Ericaceae) coffee berries (Rubiaceae) (also described as drupes)[8] Gooseberry and currant (Ribes spp.; Grossulariaceae), red, black, and white types Aubergine/Eggplant, tomato, goji berries (wolfberry) and other species of the family Solanaceae Elderberry (Sambucus niger; Adoxaceae) Indian gooseberry (Phyllanthus emblica) (Phyllanthaceae) Garcinia gummi-gutta, Garcinia mangostana (mangosteen) and Garcinia indica in the family Clusiaceae Sapodilla (Manilkara zapota), Sapotaceae Grape, Vitis vinifera in the family Vitaceae Honeysuckle: the berries of some species are edible and are called honeyberries, but others are poisonous (Lonicera spp.; Caprifoliaceae) Persimmon (Ebenaceae) Pumpkin, cucumber and watermelon in the family Cucurbitaceae Modified berries[edit] Cross-section of a cucumber pepo (Cucumis sativus) "True berries", or "baccae", may also be required to have a thin outer skin, not self-supporting when removed from the berry. This distinguishes, for example, a Vaccinium or Solanum berry from an Adansonia (baobab) amphisarca, which has a dry, more rigid and self-supporting skin.[9] The fruit of citrus, such as the orange, kumquat and lemon, is a berry with a thick rind and a very juicy interior divided into segments by septae, that is given the special name "hesperidium".[9] A specialized term, pepo, is also used for fruits of the gourd family Cucurbitaceae, which are modified to have a hard outer rind, but are not internally divided by septae.[9] The fruits of Passiflora (passion fruit) and Carica (papaya) are sometimes also considered pepos.[9] Berries that develop from an inferior ovary are sometimes termed epigynous berries or false berries, as opposed to true berries, which develop from a superior ovary. In epigynous berries, the berry includes tissue derived from parts of the flower besides the ovary. The floral tube, formed from the basal part of the sepals, petals and stamens can become fleshy at maturity and is united with the ovary to form the fruit. Common fruits that are sometimes classified as epigynous berries include bananas, coffee, members of the genus Vaccinium (e.g., cranberries and blueberries), and members of the family Cucurbitaceae (gourds, cucumbers, melons and squash).[10]


Fruits not botanical berries[edit] Several types of common "berries" Sloes (fruits of Prunus spinosa) Avocados Serviceberries (Amelanchier ovalis) Ripe mulberry (fruit of Morus nigra) In the first image, only the blueberry is botanically a berry: blackberries are aggregate fruit composed of many drupelets, and strawberries are aggregate accessory fruits. Sloes, the fruits of Prunus spinosa, are drupes. Avocado fruit are described as either drupes or berries. Serviceberries, fruits of Amelanchier species, are pomes. Mulberries, the fruits of Morus nigra, are multiple fruits. Many fruits commonly referred to as berries are not actual berries by the scientific definition, but fall into one of the following categories: Drupes[edit] Main article: Drupe Drupes are fleshy fruits produced from a (usually) single-seeded ovary with a hard woody layer (called the endocarp) surrounding the seed. Familiar examples include the stonefruits of Prunus genus (peaches, plums and cherries), olives, coconut and bayberry. Persea species. Some definitions make the mere presence of an internally differentiated endocarp the defining feature of a drupe;[9] others qualify the nature of the endocarp required in a drupe, e.g. defining berries to have endocarp less than 2 mm thick.[4] The term "drupaceous" is used of fruits that have the general structure and texture of a drupe,[11] without necessarily meeting the full definition. Other drupe-like fruits with a single seed that lack the stony endocarp include sea-buckthorn (Hippophae rhamnoides, Elaeagnaceae), which is an achene, surrounded by a swollen hypanthium that provides the fleshy layer.[12] Fruits of Coffea species are described as either drupes or berries.[8] Pomes[edit] Main article: Pome The pome fruits produced by plants in subtribe Pyrinae of family Rosaceae, such as apples and pears, have a structure (the core) in which tough tissue clearly separates the seeds from the outer softer pericarp.[13] However, some of the smaller pomes are sometimes referred to as berries. Amelanchier pomes become so soft at maturity that they resemble a blueberry and are known as Juneberries, serviceberries or Saskatoon berries.[14] Aggregate fruits[edit] Main article: Aggregate fruit Aggregate or compound fruits contain seeds from different ovaries of a single flower, with the individual "fruitlets" joined together at maturity to form the complete fruit.[15] Examples of aggregate fruits commonly called "berries" include members of the Rubus genus, such as blackberry and raspberry.[16] Other large aggregate fruits, such as soursop (Annona muricata),[17] are not usually called "berries", although some sources do use this term.[18] Multiple fruits[edit] Main article: Multiple fruit Multiple fruits are the fruits of two or more multiple flowers that are merged or packed closely together.[19] The mulberry is a berry-like example of a multiple fruit; it develops from a cluster of tiny separate flowers that become compressed as they develop into fruit.[20] Accessory fruits[edit] Main article: Accessory fruit In accessory fruits, the edible part is not generated by the ovary. Berry-like examples include: Strawberry - the aggregate of seed-like achenes is actually the "fruit", derived from an aggregate of ovaries, and the fleshy part develops from the receptacle. Mock strawberry, Duchesnea indica - structured just like a strawberry Sea grape (Coccoloba uvifera; Polygonaceae) - the fruit is a dry capsule surrounded by fleshy calyx Eastern teaberry (Gaultheria procumbens) - the fruit is a dry capsule surrounded by fleshy calyx


Berry-like conifer seed cones[edit] Yew "berries" are female conifer cones. The female seed cones of some conifers have fleshy and merged scales, giving them a berry-like appearance. Juniper "berries" (family Cupressaceae), in particular those of Juniperus communis, are used to flavour gin. The seed cones of species in the Podocarpaceae and Taxaceae families have a bright colour when fully developed, increasing the resemblance to true berries. The "berries" of yews (Taxus species) consist of a female seed cone with which develops a fleshy red aril partially enclosing the poisonous seed.


History of terminology[edit] Some fruits classified as bacca (berries) by Gaertner (De Fructibus et Seminibus Plantarum, Tab. 28) The Latin word baca or bacca (plural baccae) was originally used for "any small round fruit".[21] Andrea Caesalpinus (1519–1603) classified plants into trees and herbs, further dividing them by properties of their flowers and fruit. He did not make the modern distinction between "fruits" and "seeds", calling hard structures like nuts semina or seeds. A fleshy fruit was called a pericarpium. For Caesalpinus, a true bacca or berry was a pericarpium derived from a flower with a superior ovary; one derived from a flower with an inferior ovary was called a pomum.[22] In 1751, Carl Linnaeus wrote Philosophia Botanica, considered to be the first textbook of descriptive systematic botany.[23] He used eight different terms for fruits, one of which was bacca or berry, distinguished from other types of fruit such as drupa (drupe) and pomum (pome).[24] A bacca was defined as "pericarpium farctum evalve, semina ceteroquin nuda continens", meaning "unvalved solid pericarp, containing otherwise naked seeds".[25] The adjective "farctus" here has the sense of "solid with tissue softer than the outside; stuffed".[26] A berry or bacca was distinguished from a drupe and a pome, both of which also had an unvalved solid pericarp; a drupe also contained a nut (nux) and a pome a capsule (capsula), rather than the berry's naked seeds.[25] Linnaeus' use of bacca and pomum was thus significantly different from that of Caesalpinus. Botanists continue to differ on how fruit should be classified.[24] Joseph Gaertner published a two-volume work, De Fructibus et Seminibus Plantarum (on the fruits and seeds of plants) between 1788 and 1792. In addition to Linnaeus' eight terms, he introduced seven more, including pepo for the berry-like fruits of cucurbits.[24] A pepo was distinguished by being a fleshy berry with the seeds distant from the axis, and so nearer the fruit wall[27] (i.e. by having "parietal placentation" in modern terminology). Nicaise Auguste Desvaux in 1813 used the terms hesperidium and amphisarca as further subdivisions of berries.[24] A hesperidium, called by others bacca corticata (berry with a cortex), had separate internal compartments ("loges" in the original French) and a separable membraneous epicarp or skin. An amphisarca was described as woody on the outside and fleshy on the inside.[28] "Hesperidium" remains in general use, but "amphisarca" is rarely used.[24] There remains no universally agreed system of classification for fruits, and there continues to be "confusion over classification of fruit types and the definitions given to fruit terms".[24]


Evolution and phylogenetic significance[edit] Flowers and berries of Cestrum tomentosum By definition, berries have a fleshy, indehiscent pericarp, as opposed to a dry, dehiscent pericarp. Fossils show that early flowering plants had dry fruits; fleshy fruits, such as berries or drupes, appeared only towards the end of the Cretaceous Period or the beginning of the Paleogene Period, about 66 million years ago. The increasing importance of seed dispersal by fruit-eating vertebrates, both mammals and birds, may have driven the evolution of fleshy fruits. Alternatively, the causal direction may be the other way round. Large fleshy fruits are associated with moist habitats with closed tree canopies, where wind dispersal of dry fruits is less effective. Such habitats were increasingly common in the Paleogene and the associated change in fruit type may have led to the evolution of fruit eating in mammals and birds.[29] Fruit type has been considered to be a useful character in classification and in understanding the phylogeny of plants.[30] The evolution of fruits with a berry-like pericarp has been studied in a wide range of flowering plant families. Repeated transitions between fleshy and dry pericarps have been demonstrated regularly. One well-studied family is the Solanaceae, because of the commercial importance of fruit such as tomatoes, bell peppers, and eggplants or aubergines. Capsules, which are dry dehiscent fruits, appear to be the original form of the fruit in the earliest diverging members of the family. Berries have then evolved at least three times: in Cestrum, Duboisia, and in the subfamily Solanoideae. Detailed anatomical and developmental studies have shown that the berries of Cestrum and those of the Solanoideae are significantly different; for example, expansion of the fruit during development involves cell divisions in the mesocarp in Solanoideae berries, but not in Cestrum berries.[31] When fruits described as berries were studied in the family Melastomaceae, they were found to be highly variable in structure, some being soft with an endocarp that soon broke down, others having a hard, persistent endocarp, even woody in some species.[30] Fruits classified as berries are thus not necessarily homologous, with the fleshy part being derived from different parts of the ovary, and with other structural and developmental differences. The presence or absence of berries is not a reliable guide to phylogeny.[29] Indeed, fruit type in general has proved to be an unreliable guide to flowering plant relationships.[30]


Uses[edit] Culinary[edit] A type of sapote fruit displayed for sale (Quararibea cordata) Bottle gourd or calabash used to contain palm wine in Bandundu Province, Democratic Republic of the Congo Mandarins, here served in a Hong Kong restaurant, are among the oldest cultivated citrus fruits. Berries, defined loosely, have been valuable as a food source to humans since prior to the start of agriculture, and remain among the primary food sources of other primates. Botanically defined berries with culinary uses include: Berries in the strictest sense: including bananas and plantains, blueberries, cranberries, the seeds of coffee berries, gooseberries, red-, black- and white currants, tomatoes, grapes and peppers (Capsicum fruits) Hesperidia: citrus fruits, including oranges, lemons and limes Pepos: cucurbits, including squashes, cucumbers, melons and watermelons Some berries are brightly coloured, due to plant pigments such as anthocyanins and other flavonoids. These pigments are localized mainly in the outer surface and the seeds.[32] Such pigments have antioxidant properties in vitro,[33] but there is no reliable evidence that they have antioxidant or any other useful functions within the human body. Consequently, it is not permitted to claim that foods containing plant pigments have antioxidant health value on product labels in the United States or Europe.[34][35] Some spices are prepared from berries. Allspice is made from the dried berries of Pimenta dioica.[36] The fruits (berries) of different cultivars of Capsicum annuum are used to make paprika (mildly hot), chili pepper (hot) and cayenne pepper (very hot).[37] Others[edit] Pepos, characterized by a hard outer rind, have also been used as containers by removing the inner flesh and seeds and then drying the remaining exocarp. The English name of Lagenaria siceraria, "bottle gourd", reflects its use as a liquid container.[38] Some true berries have also been used as a source of dyes. In Hawaii, these included berries from a species of Dianella, used to produce blue, and berries from black nightshade (Solanum americanum), used to produce green.[39] History[edit] Cucurbit berries or pepos, particularly from Cucurbita and Lagenaria, are the earliest plants known to be domesticated – before 9,000–10,000 BP in the Americas, and probably by 12,000–13,000 BP in Asia.[38] Peppers were domesticated in Mesoamerica by 8,000 BP.[40][41] Many other early cultivated plants were also berries by the strict botanical definition, including grapes, domesticated by 8,000 BP[42][43] and known to have been used in wine production by 6,000 BP.[44] Bananas were first domesticated in Papua New Guinea and Southeast Asia. Archaeological and palaeoenvironmental evidence at Kuk Swamp in the Western Highlands Province of Papua New Guinea suggests that banana cultivation there goes back to at least 7,000 BP, and possibly to 10,000 BP.[45][46] The history of cultivated citrus fruit remains unclear, although some recent research suggests a possible origin in Papuasia rather than continental south-east Asia. Chinese documents show that mandarins and pomelos were established in cultivation there by around 4,200 BP.[47] Commercial production[edit] Four banana and plantain cultivars Watermelon Grapes According to FAOSTAT data, in 2013 four of the five top fruit crops in terms of world production by weight were botanical berries. The other was a pome (apples).[Note 1] Worldwide fruit production in 2013 Name 1000s of tonnes Fruit type Bananas & plantains 144,592 Berry Citrus fruit† 135,761 Berry (hesperidium) Watermelons 109,279 Berry (pepo) Apples 80,823 Pome Grapes 77,181 Berry †Citrus fruit includes, but is not limited to, oranges, lemons, limes, grapefruit (including pomelos), tangerines, mandarins, clementines and satsumas. Oranges make up 53% of the total. According to FAOSTAT, in 2001, bananas (including plantains) and citrus comprised over 25% by value of the world's exported fruits and vegetables, citrus fruits being more valuable than bananas.[49] Export quantities of fruit are not entirely comparable with production quantities, since slightly different categories are used. The top five fruit exports by weight in 2012 are shown in the table below. The top two places are again occupied by bananas and citrus.[Note 2] Worldwide fruit export in 2012 Name 1000s of tonnes Fruit type Bananas & plantains 19,725 Berry Citrus fruit† 15,262 Berry (hesperidium) Apples 8,271 Pome Prepared fruit‡ 7,120 – Grapes 4,051 Berry †Citrus fruit includes oranges, lemons, limes, grapefruit (including pomelos), tangerines, mandarins, clementines and satsumas. Oranges make up 43% of the total. ‡Prepared fruit here is "fruit, nuts and peel, including frozen, prepared or preserved, jam, paste, marmalade, purée and cooked fruits, other than those listed separately".[50]


See also[edit] List of culinary fruits List of inedible fruits


Notes[edit] ^ Production Quantity data for 2013 for the World total for all items was downloaded to a spreadsheet from FAOSTAT.[48] Items coded 486 to 626, corresponding to fruit, were retained. Data for bananas and plantains were combined, as these are not distinguished by all countries. Data for all citrus fruits were also combined, since some countries provide data under a general heading of "citrus fruit". The resulting table was then sorted and the top five entries used. ^ Export Quantity data for 2012 for the World total for all items was downloaded to a spreadsheet from FAOSTAT.[48] As for production, items coded 486 to 626 (but now excluding 564 wine, not present in the production data) were retained. Data for bananas and plantains were combined, as was data for all citrus fruits. The resulting table was then sorted and the top five entries used.


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External links[edit] Look up Berry in Wiktionary, the free dictionary. Wikimedia Commons has media related to Fruit by taxon. v t e Types of fruits Types of fruits Achene Berry (modified berries: Hesperidium Pepo) Capsule Caryopsis Drupe Follicle Legume Loment Nut Pome Samara Schizocarp Silique Syconium Categories of fruits Accessory fruit Simple fruit Compound fruit Aggregate fruit Multiple fruit Dehiscent fruit Function Diaspore Drift fruit v t e Botany History of botany Subdisciplines Plant systematics Ethnobotany Paleobotany Plant anatomy Plant ecology Phytogeography Geobotany Flora Phytochemistry Plant pathology Bryology Phycology Floristics Dendrology Plant groups Algae Archaeplastida Bryophyte Non-vascular plants Vascular plants Spermatophytes Pteridophyte Gymnosperm Angiosperm Plant morphology (glossary) Plant cells Cell wall Phragmoplast Plastid Plasmodesma Vacuole Tissues Meristem Vascular tissue Vascular bundle Ground tissue Mesophyll Cork Wood Storage organs Vegetative Root Rhizoid Bulb Rhizome Shoot Stem Leaf Petiole Cataphyll Bud Sessility Reproductive (Flower) Flower development Inflorescence Umbel Raceme Bract Pedicellate Flower Whorl Floral symmetry Floral diagram Floral formula Receptacle Hypanthium (Floral cup) Perianth Tepal Petal Sepal Sporophyll Gynoecium Ovary Ovule Stigma Archegonium Androecium Stamen Staminode Pollen Tapetum Gynandrium Gametophyte Sporophyte Plant embryo Fruit Fruit anatomy Berry Capsule Seed Seed dispersal Endosperm Surface structures Epicuticular wax Plant cuticle Epidermis Stoma Nectary Trichome Prickle Plant physiology Materials Nutrition Photosynthesis Chlorophyll Plant hormone Transpiration Turgor pressure Bulk flow Aleurone Phytomelanin Sugar Sap Starch Cellulose Plant growth and habit Secondary growth Woody plants Herbaceous plants Habit Vines Lianas Shrubs Subshrubs Trees Succulent plants Reproduction Evolution Ecology Alternation of generations Sporangium Spore Microsporangia Microspore Megasporangium Megaspore Pollination Pollinators Pollen tube Double fertilization Germination Evolutionary development Evolutionary history timeline Hardiness zone Plant taxonomy History of plant systematics Herbarium Biological classification Botanical nomenclature Botanical name Correct name Author citation International Code of Nomenclature for algae, fungi, and plants (ICN) - for Cultivated Plants (ICNCP) Taxonomic rank International Association for Plant Taxonomy (IAPT) Plant taxonomy systems Cultivated plant taxonomy Citrus taxonomy cultigen cultivar Group grex Practice Agronomy Floriculture Forestry Horticulture Lists Related topics Botanical terms Botanists by author abbreviation Botanical expedition Category Portal WikiProject Plants portal Food portal Retrieved from "https://en.wikipedia.org/w/index.php?title=Berry_(botany)&oldid=802559491" Categories: BerriesFruit morphologyHidden categories: CS1 Latin-language sources (la)Articles containing Latin-language textCS1 French-language sources (fr)Use dmy dates from August 2015Articles containing French-language textCommons category with local link different than on Wikidata


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ProvinceDemocratic Republic Of The CongoEnlargeHong KongCapsicumPlant PigmentAnthocyaninFlavonoidSeedAntioxidantIn VitroSpiceAllspiceCultivarCapsicum AnnuumCalabashDyeHawaiiDianella (plant)Solanum AmericanumDomesticationPapua New GuineaSoutheast AsiaArchaeologyKuk SwampWestern Highlands ProvinceCitrusPapuasiaMandarin OrangePomeloEnlargeEnlargeEnlargeFood And Agriculture Organization Corporate Statistical DatabaseList Of Culinary FruitsList Of Inedible FruitsInternational Standard Book NumberSpecial:BookSources/978-1-84246-422-9Digital Object IdentifierInternational Standard Book NumberSpecial:BookSources/978-81-7133-896-2Clive StaceInternational Standard Book NumberSpecial:BookSources/978-0-521-70772-5International Standard Book NumberSpecial:BookSources/978-0-19-217674-5Google BooksInternational Standard Book NumberSpecial:BookSources/978-0-7153-1643-6Digital Object IdentifierPubMed IdentifierDigital Object IdentifierDigital Object IdentifierPubMed IdentifierJournal Of Agricultural And Food ChemistryDigital Object IdentifierPubMed IdentifierDigital Object IdentifierInternational Standard Book NumberSpecial:BookSources/978-0-85199-605-9Digital Object IdentifierPubMed CentralPubMed IdentifierInternational Standard Book NumberSpecial:BookSources/978-0-8248-0698-9Digital Object IdentifierDigital Object IdentifierPubMed CentralPubMed IdentifierDigital Object IdentifierDigital Object IdentifierPubMed IdentifierDigital Object IdentifierTemplate:FruitsTemplate Talk:FruitsFruitAcheneHesperidiumBerry (botany)Capsule (fruit)CaryopsisDrupeFollicle (fruit)LegumeLomentNut (fruit)PomeSamara (fruit)SchizocarpSiliqueSyconiumAccessory FruitFruitCompound FruitAggregate FruitMultiple FruitDehiscence (botany)Diaspore (botany)Drift SeedTemplate:BotanyTemplate Talk:BotanyBotanyHistory Of BotanyBranches Of BotanyHistory Of Plant SystematicsEthnobotanyPaleobotanyPlant AnatomyPlant EcologyPhytogeographyGeobotanical ProspectingFloraPhytochemistryPlant PathologyBryologyPhycologyFloristicsDendrologyPlantAlgaeArchaeplastidaBryophyteNon-vascular PlantVascular PlantSpermatophytePteridophyteGymnospermFlowering PlantPlant MorphologyGlossary Of Plant MorphologyPlant CellCell WallPhragmoplastPlastidPlasmodesmaVacuoleTissue (biology)MeristemVascular TissueVascular BundleGround TissueLeafCork CambiumWoodStorage OrganRootRhizoidBulbRhizomeShootPlant StemLeafPetiole (botany)CataphyllBudSessility (botany)Plant Reproductive MorphologyABC Model Of Flower DevelopmentInflorescenceUmbelRacemeBractPedicel (botany)FlowerWhorl (botany)Floral SymmetryFloral DiagramFloral FormulaReceptacle (botany)HypanthiumPerianthTepalPetalSepalSporophyllGynoeciumOvary (botany)OvuleStigma (botany)ArchegoniumStamenStamenStaminodePollenTapetum (botany)Column (botany)GametophyteSporophyteEmbryoFruitFruit AnatomyCapsule (fruit)SeedSeed DispersalEndospermEpicuticular WaxPlant CuticleEpidermis (botany)StomaNectarTrichomeThorns, Spines, And PricklesPlant PhysiologyPlant NutritionPhotosynthesisChlorophyllPlant HormoneTranspirationTurgor PressureBulk MovementAleuronePhytomelaninSugarSapStarchCelluloseSecondary GrowthWoody PlantHerbaceous PlantHabit (biology)VineLianaShrubSubshrubTreeSucculent PlantPlant ReproductionPlant EvolutionPlant EcologyAlternation Of GenerationsSporangiumSporeMicrosporangiaMicrosporeSporangiumMegasporePollinationPollinatorPollen TubeDouble FertilizationGerminationPlant Evolutionary Developmental BiologyEvolutionary History Of PlantsTimeline Of Plant EvolutionHardiness ZonePlant TaxonomyHistory Of Plant SystematicsHerbariumTaxonomy (biology)Botanical NomenclatureBotanical NameCorrect NameAuthor Citation (botany)International Code Of Nomenclature For Algae, Fungi, And PlantsInternational Code Of Nomenclature For Cultivated PlantsTaxonomic RankInternational Association For Plant TaxonomyList Of Systems Of Plant TaxonomyCultivated Plant TaxonomyCitrus TaxonomyCultigenCultivarCultivar GroupGrex 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